Bark and Ambrosia Beetles of

SYNONYMY

Taxonomic Comments.   For the present the treatment of this species follows Lanier et. al (1991) and treats all Central Mexican and Central American records as I. apache Lanier, even though no attempt has been made to identify specimens based on the characters given in that publication.

• Ips calligraphus Auctorum & not Germar 0. various
• Ips apache Lanier 1991. Can. Ent. 123: 1118

DISTRIBUTION.

Numbers in parentheses after each geographic unit are the number of distinct collection events in the database for that unit. For exotic species generally only countries are listed for localities outside the New World. For further information on published sources of distribution, check the REFERENCES section.

• Central America (13)
  Belize (1): unknown (1); Guatemala (2): Baja Verapaz (1), Chichicastenango (1); Honduras (4): Comayagua (1), Cortés (1), Cortes (1), Olancho (1); Nicaragua (2): Estelí (1), unknown (1); Panamá (4): Chiriquí (3), Panamá (1);
• North America (25)
  (1): no state / prov. data (1); México (12): Chiapas (2), Guerrero (2), Hidalgo (1), Jalisco (2), México (3), Michoacán (2); Mexico (1): Michoacán (1); México (9): Morelos (2), Oaxaca (4), Querétaro (1), Sinaloa (1), Veracruz (1); United States (2): Arizona (2);

HOSTS

Numbers in parentheses after each host family, genus, or collection method are the number of distinct collection events in the database for that host or method.

• No host information (12);
• Pinaceae (25): Pinus (25): no species data (1), devoniana (1), engelmanni (2), michoacana (1), montezumae (2), oocarpa (4), pseudostrobus (3), sp. (10), tenuifolia (1);
• trap (1) Malaise (1): no details (1);

REFERENCES

The following are important recent monographs, catalogs, and supplements to catalogs that refer to this species. The specific page on which the reference is made is shown in pink at the end of the reference. In the case of Wood (1982) and Wood & Bright (1992) clicking on the reference page will link to a digital version of the work in question.

• Wood, S.L., Bright,D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index. Great Basin Nat. Mem. 13:1-1553 (vol. A, B). [491]

The following are references from which host and distribution data have been input into the database. If one of the above monographs or catalogs also appears in this list, it means that most relevant collection event data have been included.

1. . . .
2. Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986. Coleópteros Scolytidae y Platypodidae asociados con las comunidades vegetales de clima templado y frío en el estado de Morelos. Acta Zool. Mex. 17: 1-58.
3. Cognato, A.I.; Sperling, F.A.H. 2000. Phylogeny of Ips DeGeer species (Coleoptera: Scolytidae) inferred from mitochondrial cytochrome oxidase I DNA sequence. Mol. Phylogenet. Evol. 14 (3): 445-460.
4. Hopping, G.R. 1965. The North American species in Group X of Ips DeGeer (Coleoptera: Scolytidae). Can. Entomol. 97(8): 803-809.
5. Morales M., R.; Castrejón A., J.E.; Sánchez R., J.C.; Ballesteros T., J.; Jaimes Y., N.; Fuentes G., L.T. 2023. Ips apache Infestation of Pinus devoniana (= Pinus michoacana) in Southern Jalisco, Mexico. Southwest. Entomol. 48 (3): 721-724.
6. Smith, S.M.; Cognato, A.I. 2009. Occurrence of Ips apache Lanier (Coleoptera: Curculionidae: Scolytinae) in Panama. Coleop. Bull 63(4): 452-453.
7. Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6: 1-1356.
8. Wood, S.L.; Bright, D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index (Volumes A,B). Great Basin Nat. Mem. 13: 1-1553.

Maps automatically open at the center of the plotted points and the scale is set to encompass all map points. Maps can be resized (scale bar at upper left) and the center moved (place cursor over map and drag) to see other parts of the distribution of the species. Clicking on a map marker will pull up collection event data and a literature citation if present. If any errors are found, please refer to the "series code" which is a unique identifier for a database record in any communications). Coordinates have not been entered for all collection records. Localities outside the New World are not plotted, even though they are listed in the distribution summary and in the table of records.

No.	Collection data	Publication
Central America
1->	Belize, unknown, Belize (country)	[unknown () (Hopping, G.R. 1965: 803)]
2->	Guatemala, Baja Verapaz, San Jerónimo	[unknown () (Hopping, G.R. 1965: 803)]
3->	Guatemala, Chichicastenango, Chichicastenango; 23/VII/1980	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 235)]
4->	Honduras, Comayagua, Los Planes, 2.8 km NNE; 26/V/2002; Turnbow, R.H.	[RHTC (1) (TH Atkinson-2008, unpublished)]
5->	Honduras, Cortés, Agua Azul vic.; 30/V/1993; Skillman, F.W.	[FSCA (1) (TH Atkinson-2013, unpublished)]
6->	Honduras, Cortes, Parque Nacional Cusuco; 30/V/2010; LLAMA Project; trap: Malaise	[SEMC (6) (TH Atkinson-2020, unpublished)]
7->	Honduras, Olancho, La Unión, 10 km N; 29/XI/1995; Turnbow, R.H.; Pinaceae: Pinus sp.	[FSCA (6) (TH Atkinson-2007, unpublished)] [RHTC (17) (TH Atkinson-2008, unpublished)]
8->	Nicaragua, Estelí, La Almaciguera; 14/III/2010; Atkinson, T.H.; Pinaceae: Pinus oocarpa	[UTIC (5) (TH Atkinson-2010, unpublished)]
9->	Nicaragua, unknown, Nicaragua (country)	[unknown () (Wood, S.L. 1982: 697)]
10->	Panamá, Chiriquí, Bajo Mono; VIII/2008; Smith, S.M.; Pinaceae: Pinus sp.	[MSUC (1) (Smith, S.M.; Cognato, A.I. 2009: 453)]
11->	Panamá, Chiriquí, Fortuna; VIII/2008; Smith, S.M.; Pinaceae: Pinus sp.	[MSUC (1) (Smith, S.M.; Cognato, A.I. 2009: 452)]
12->	Panamá, Chiriquí, Sta. Clara; VIII/2008; Smith, S.M.; Pinaceae: Pinus sp.	[MSUC (1) (Smith, S.M.; Cognato, A.I. 2009: 452)]
13->	Panamá, Panamá, Cerro Jefe; VIII/2008; Smith, S.M.; Pinaceae: Pinus sp.	[MSUC (1) (Smith, S.M.; Cognato, A.I. 2009: 452)]
North America
14->	, no state / prov. data; Pinaceae: Pinus michoacana	[unknown (1) (Wood, S.L. 1982: 697)]
15->	México, Chiapas, Altamirano	[unknown () (Hopping, G.R. 1965: 803)]
16->	México, Chiapas, Angel Corzo; 5/VI/1985; Bermúdez, N.; Pinaceae: Pinus tenuifolia	[unknown (1) (Atkinson, T.H.; Equihua M., A. 1988: 94)]
17->	México, Guerrero, Chilapa; 23/II/1982; Atkinson, T.H.; Equihua M., A.; Pinaceae: Pinus oocarpa	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 235)]
18->	México, Guerrero, Taxco; 22/II/1982; Atkinson, T.H.; Equihua M., A.; Pinaceae: Pinus sp.	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 235)]
19->	México, Hidalgo, Zimapan	[unknown () (Hopping, G.R. 1965: 803)]
20->	México, Jalisco, El Tuito; 6/III/1982; Equihua M., A.; Pinaceae: Pinus sp.	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 235)]
21->	México, Jalisco, Pihuamo; VIII/2022; Pinaceae: Pinus devoniana	[BCUC (1) (Morales M., R.; Castrejón A., J.E.; Sánchez R., J.C.; Ballesteros T., J.; Jaimes Y., N.; Fuentes G., L.T. 2023: 721)]
22->	México, México, Temascaltepec; 17/VIII/1960; Pinaceae: Pinus oocarpa	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 235)]
23->	México, México, Temascaltepec	[unknown () (Hopping, G.R. 1965: 803)]
24->	México, México, Valle de Bravo; 9/III/1962	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 235)]
25->	México, Michoacán, Ario de Rosales; 31/X/1980; Atkinson, T.H.; Equihua M., A.; Pinaceae: Pinus pseudostrobus	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 235)]
26->	México, Michoacán, Ario de Rosales; 31/X/1980; Atkinson, T.H.; Equihua M., A.; Pinaceae: Pinus pseudostrobus	[FSCA (4) (TH Atkinson-1981, unpublished)]
27->	Mexico, Michoacán, Barranca del Cupatitzio; 1996; Storer, A.	[EMEC (1) (Cognato, A.I.; Sperling, F.A.H. 2000: 457)]
28->	México, Morelos, Chamilpa; Pinaceae: Pinus montezumae	[UAEM () (Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986: 31)]
29->	México, Morelos, Chamilpa; Pinaceae: Pinus montezumae	[UAEM () (Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986: 31)]
30->	México, Oaxaca, Benito Juárez; 5/II/1984; Equihua M., A.; Pinaceae: Pinus	[CEAM () (Atkinson, T.H.; Equihua M., A. 1988: 92)]
31->	México, Oaxaca, Miahuatlan, 5 km N; 18/XI/1989; Baranowski, R.M.; Pinaceae: Pinus sp.	[ZMLU (1) (TH Atkinson-2017, unpublished)]
32->	México, Oaxaca, Sta. Ma. Chimalapa; 9/II/1984; Equihua M., A.; Pinaceae: Pinus oocarpa	[CEAM () (Atkinson, T.H.; Equihua M., A. 1988: 92)]
33->	México, Oaxaca, Textlihuaca, 19 km NW. carr. 135; 22/X/2011; Atkinson, T.H.; Pinaceae: Pinus sp.	[UTIC (1) (TH Atkinson-2012, unpublished)]
34->	México, Querétaro, Landa de Matamoros, 18 mi E; 18/VI/1983; O'Brien, C.W.; O'Brien, L.B.; Marshall, G.B.; Pinaceae: Pinus sp.	[FSCA (22) (TH Atkinson-2007, unpublished)]
35->	México, Sinaloa, Loberas, 9 km NE ; 30/IX/1990; Turnbow, R.H.	[RHTC (1) (TH Atkinson-1993, unpublished)]
36->	México, Veracruz, El Escolín; 16/XII/1983; Noguera M., F.A.; Pinaceae: Pinus pseudostrobus	[CEAM () (Atkinson, T.H.; Equihua M., A. 1988: 92)]
37->	United States, Arizona, Portal, 10 mi NW, Coronado N.F. Rd. 42; 6/IX/1996; Cognato, A.I.; Pinaceae: Pinus engelmanni	[EMEC (1) (Cognato, A.I.; Sperling, F.A.H. 2000: 457)]
38->	United States, Arizona, Portal, 7 mi W; VIII/1972; Lanier, G.N.; Pinaceae: Pinus engelmanni	[USNM (1: holotype) (Wood, S.L.; Bright, D.E. 1992: 491) (TH Atkinson-2018, unpublished)]