Bark and Ambrosia Beetles of

SYNONYMY

Taxonomic Comments.   Some specimens reported from the southeastern United States may actually be Hypothenemus subterrestris Johnson, Atkinson & Hulcr 2016.

• Hypothenemus californicus Hopkins 1915. U.S. Dep. Agric. Tech. Bull. (99): 19
• Hypothenemus californicus triciti Hopkins 1915. U.S. Dep. Agric. Tech. Bull. (99): 19
• Hypothenemus californicus californicus Hopkins 1915. U.S. Dep. Agric. Tech. Bull. (99): 19
• Hypothenemus tritici Hopkins 1915. U.S. Dep. Agric. Tech. Bull. (99): 19
• Hypothenemus thoracicus Hopkins 1916. Scolytidae (in Blatchley & Leng 1916): 598
• Stephanoderes zeae Schedl 1973. Pap. Avulsos Depto. Zool. 26: 169

DISTRIBUTION.

Numbers in parentheses after each geographic unit are the number of distinct collection events in the database for that unit. For exotic species generally only countries are listed for localities outside the New World. For further information on published sources of distribution, check the REFERENCES section.

Distribution Comments.   Widely distributed but seldom collected in tropical and subtropical regions of the Americas. Southeastern U.S. and southern California to Brazil.

• Africa (1)
  Liberia (1): no state / prov. data (1);
• Central America (1)
  Belize (1): Orange Walk (1);
• Caribbean (1)
  Lesser Antilles (1): St. Lucia (1);
• Eurasia (2)
  Israel (1): no state / prov. data (1); Korea (1): no state / prov. data (1);
• North America (140)
  México (39): Guerrero (1), Jalisco (2), Michoacán (2), Morelos (28), Oaxaca (1), Sonora (1), Tamaulipas (1), Veracruz (3); United States (101): Alabama (6), Arkansas (1), California (8), District of Columbia (1), Florida (16), Georgia (10), Illinois (1), Indiana (1), Kansas (2), Kentucky (1), Louisiana (5), Maryland (1), Mississippi (4), Missouri (2), New Jersey (1), North Carolina (1), Oklahoma (6), South Carolina (8), Tennessee (1), Texas (23), Virginia (2);
• South America (9)
  Brazil (4): Bahia (1), Mato Grosso do Sul (1), Paraiba (1), São Paulo (1); Ecuador (5): Galápagos (5);
• UNK (17)
  (17): no state / prov. data (17);

HOSTS

Numbers in parentheses after each host family, genus, or collection method are the number of distinct collection events in the database for that host or method.

• No host information (79);
• Amaranthaceae (1): (1): no species data (1);
• Anacardiaceae (2): Mangifera (1): indica (1); Toxicodendron (1): radicans (1);
• Apocynaceae (2): Funastrum (1): clausum (1); Thevetia (1): ovata (1);
• Arecaceae (1): Sabal (1): pomos (1);
• Combretaceae (1): Quisqualis (1): indica (1);
• Compositae (3): Bidens (1): pilosa (1); Encelia (1): farinosa (1); Iva (1): imbricata (1);
• Convolvulaceae (2): Ipomoea (2): cathartica (1), littoralis (1);
• Cucurbitaceae (7): (5): no species data (5); Cucurbita (1): argyrosperma (1); Luffa (1): cylindrica (1);
• Euphorbiaceae (3): Croton (2): ciliato-glandulosum (1), sp. (1); Sapium (1): macrocarpum (1);
• Leguminosae (7): (1): no species data (1); Bauhinia (1): alba (1); Cajanus (1): cajon (1); Galactia (1): spiciformis (1); Mimosa (1): albida (1); Phaseolus (2): sp. (2);
• Liliaceae (1): Aloe (1): vera (1);
• litter (29): berlese (29): no species data (5), beech litter (1), beech, holly litter (1), beech, pine, grass litter (1), hardwood litter (1), leaf litter (9), litter hardwood forest (1), longleaf pine, oak litter (1), mixed hardwood leaf litte (2), mixed hardwood litter (1), mixed pine hardwood leaf (1), oak litter (1), pine - wiregrass litter (1), pine litter (1), sifted pine oak litter (1), sifted roadside scrub lit (1);
• Malvaceae (1): Malvastrum (1): sp. (1);
• Moraceae (5): Artocarpus (1): heterophyllus (1); Ficus (4): carica (1), elastica (1), sp. (2);
• Poaceae (4): Paspalum (1): vaginatum (1); Triticum (1): sp. (1); Uniola (1): paniculata (1); Zea (1): mays (1);
• Salicaceae (1): Salix (1): babylonica (1);
• Santalaceae (1): Phoradendron (1): sp. (1);
• Sapotaceae (1): Sideroxylon (1): capiri (1);
• Scrophulariaceae (1): Capraria (1): biflora (1);
• trap (18): flight intercept (6): no details (5), ethanol (1); Lindgren Funnel (3): no details (2), turpentine (1); pitfall (5): no details (5); window trap (4): no details (2), ethanol (1), unbaited (1);
• Verbenaceae (1) Verbena (1): sp. (1);

REFERENCES

The following are important recent monographs, catalogs, and supplements to catalogs that refer to this species. The specific page on which the reference is made is shown in pink at the end of the reference. In the case of Wood (1982) and Wood & Bright (1992) clicking on the reference page will link to a digital version of the work in question.

• Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6:1-1356. [897]. (data capture complete)
• Wood, S.L., Bright,D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index. Great Basin Nat. Mem. 13:1-1553 (vol. A, B). [912]
• Wood, S.L. 2007. Bark and ambrosia beetles of South America (Coleoptera: Scolytidae). Monte L. Bean Sci. Mus.,Provo, Utah. 900 pp. [517]. (data capture complete)

The following are references from which host and distribution data have been input into the database. If one of the above monographs or catalogs also appears in this list, it means that most relevant collection event data have been included.

1. . . .
2. Atkinson, T.H. 2019. Escarabajos descortezadores y ambrosiales (Coleoptera: Curculionidae: Scolytinae, Platypodinae) de Sonora, México (Bark and Ambrosia Beetles (Coleoptera: Curculionidae: Scolytinae, Platypodinae) of Sonora, México). Dugesiana 26(1): 41-49.
3. Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986. Scolytidae y Platypodidae (Coleoptera) asociados a selva baja caducifolia y comunidades derivadas en el estado de Morelos. Fol. Entomol. Mex. 69: 41-82.
4. Atkinson, T.H.; Riley, E.G. 2013. Atlas and Checklist of the bark and ambrosia beetles of Texas and Oklahoma (Curculionidae: Scolytinae and Platypodinae). Insecta Mundi 292: 1-46.
5. Bright, D.E., Skidmore, R.E. 2002. A Catalog of Scolytidae and Platypodidae (Coleoptera), Supplement 2 (1995-1999). NRC Research Press. Ottawa, Ontario, Canada.
6. Bright, D.E., Stark, R.W. 1973. The bark and ambrosia beetles of California. Coleoptera: Scolytidae and Platypodidae. Bull. Calif. Insect Surv. 16: 1-169.
7. Hopkins, A.D. 1915. Classification of the Cryphalinae with descriptions of new genera and species. USDA Ent. Comm. Report (99): 1-75.
8. Johnson, A.J.; Atkinson, T.H.; Hulcr, J. 2016. Two remarkable new species of Hypothenemus Westwood (Curculionidae: Scolytinae) from Southeastern USA . Zootaxa 4200(3): 417-425.
9. Weber, B.C., MacPherson, J.E. 1991. Seasonal Flight Patterns Of Scolytidae (Coleoptera) In Black-Walnut Plantations In North-Carolina And Illinois. Coleopterists Bull. 45(1): 45-56.
10. Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6: 1-1356.
11. Wood, S.L. 2007. Bark and ambrosia beetles of South America (Coleoptera: Scolytidae). Monte L. Bean Science Museum. Provo, Utah: 1-900.
12. Wood, S.L.; Bright, D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index (Volumes A,B). Great Basin Nat. Mem. 13: 1-1553.

Maps automatically open at the center of the plotted points and the scale is set to encompass all map points. Maps can be resized (scale bar at upper left) and the center moved (place cursor over map and drag) to see other parts of the distribution of the species. Clicking on a map marker will pull up collection event data and a literature citation if present. If any errors are found, please refer to the "series code" which is a unique identifier for a database record in any communications). Coordinates have not been entered for all collection records. Localities outside the New World are not plotted, even though they are listed in the distribution summary and in the table of records.

No.	Collection data	Publication
Africa
1->	Liberia, no state / prov. data	[unknown () (Wood, S.L. 1982: 897)]
Central America
2->	Belize, Orange Walk, Rio Bravo Conservation Area; 15/V/1997; Carlton, C.E.; trap: flight intercept	[LSAM (1) (TH Atkinson-2013, unpublished)]
Caribbean
3->	Lesser Antilles, St. Lucia, Sta. Lucia; Weber, N.A.	[MCZ (1) (TH Atkinson-1988, unpublished)]
Eurasia
4->	Israel, no state / prov. data	[unknown () (Wood, S.L. 1982: 897)]
5->	Korea, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 912)]
North America
6->	México, Guerrero, El Ocotito, 7.2 km NW; 7/VII/1987; Anderson, R.; litter: berlese	[TAMU (1) (TH Atkinson-1988, unpublished)]
7->	México, Jalisco, Chamela; Amaranthaceae	[CEAM () (Equihua M., A.; Atkinson, T.H. 1986: 631)]
8->	México, Jalisco, Magdalena, 7 mi W; 26/X/1982; Huber, J.	[CSCA (1) (TH Atkinson-2021, unpublished)]
9->	México, Michoacán, La Gallina; 1/XI/1980; Atkinson, T.H.; Equihua M., A.; Arecaceae: Sabal pomos	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 357)]
10->	México, Michoacán, Quiroga	[Unknown () (Wood, S.L. 1982: 897)]
11->	México, Morelos, Amador Salazar	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
12->	México, Morelos, Cañon de Lobos; Burgos S., A.	[CEAM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
13->	México, Morelos, Cañon de Lobos	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
14->	México, Morelos, Carr. a Moyotepec	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
15->	México, Morelos, Carr. Alpuyeca-Grutas, km 8; Euphorbiaceae: Croton ciliato-glandulosum	[CEAM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
16->	México, Morelos, Carr. Hornos-Chimalacatlan; Burgos S., A.; Sapotaceae: Sideroxylon capiri	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
17->	México, Morelos, Carr. Hornos-Chimalacatlan; Cucurbitaceae	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
18->	México, Morelos, Carr. Moyotepec-Temilpa	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
19->	México, Morelos, Carr. Xochimilco-Oaxtepec, km 58; Leguminosae: Phaseolus sp.	[CEAM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)] [FSCA (2) (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
20->	México, Morelos, Cerro la Herradura; Leguminosae: Phaseolus sp.	[CEAM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
21->	México, Morelos, Cerro la Herradura; Burgos S., A.; Leguminosae: Mimosa albida	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
22->	México, Morelos, Cerro la Herradura; Cucurbitaceae	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
23->	México, Morelos, Cuernavaca; Burgos S., A.; Cucurbitaceae	[CEAM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
24->	México, Morelos, Cuernavaca; Burgos S., A.; Cucurbitaceae	[CEAM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
25->	México, Morelos, Cuernavaca; Burgos S., A.; Moraceae: Ficus elastica	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
26->	México, Morelos, Cuernavaca; Burgos S., A.; Cucurbitaceae: Luffa cylindrica	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
27->	México, Morelos, Cuernavaca	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
28->	México, Morelos, Cuernavaca; Euphorbiaceae: Sapium macrocarpum	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
29->	México, Morelos, Cuernavaca	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
30->	México, Morelos, Las Piedras; Burgos S., A.; Apocynaceae: Thevetia ovata	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
31->	México, Morelos, Morelos (estado)	[Unknown () (Wood, S.L. 1982: 897)]
32->	México, Morelos, Santiago Tepetlapa; Leguminosae	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
33->	México, Morelos, Sn. Rafael Zaragoza; Burgos S., A.	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
34->	México, Morelos, Sta. Rosa; Cucurbitaceae	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
35->	México, Morelos, Tepalcingo; Moraceae: Ficus sp.	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
36->	México, Morelos, Yautepec	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
37->	México, Morelos, Yautepec	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
38->	México, Morelos, Yautepec	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 64)]
39->	México, Oaxaca, Finca Monte Carlo, above Sta. Ma. Xadani; 24/VI/2009; Atkinson, T.H.; Moraceae: Ficus sp.	[TAMU (2) (TH Atkinson-2009, unpublished)]
40->	México, Sonora, Alamos Hwy, Aduana turnoff; 31/I/2016; Atkinson, T.H.; Smith, S.M.; Cucurbitaceae: Cucurbita argyrosperma	[UTIC (1) (Atkinson, T.H. 2019: 44)]
41->	México, Tamaulipas, Victoria	[Unknown () (Wood, S.L. 1982: 897)]
42->	México, Veracruz, El Castillo; Santalaceae: Phoradendron sp.	[CEAM (8) (Noguera M., F.A., Atkinson, T.H. 1990: 464)]
43->	México, Veracruz, Orizaba, 5 km E	[Unknown () (Wood, S.L. 1982: 897)]
44->	México, Veracruz, Xalapa; 1/IV/1984; Noguera M., F.A.; Anacardiaceae: Toxicodendron radicans	[TAMU (14) (TH Atkinson-1987, unpublished)]
45->	United States, Alabama, Brewton, 17.5 mi E; 5/II/2003; O'Brien, C.W.; Haseeb, M.; litter: berlese beech, holly litter	[FSCA (1) (TH Atkinson-2007, unpublished)]
46->	United States, Alabama, Co. Rd. 4, 0.5 mi E jct. 16; 9/II/2002; O'Brien, C.W.; Turnbow, R.H.; litter: berlese mixed hardwood leaf litte	[RHTC (2) (TH Atkinson-2008, unpublished)]
47->	United States, Alabama, Hwy 189, 1,5 mi N jct co rd 337; 10/II/2002; O'Brien, C.W.; Turnbow, R.H.; litter: berlese sifted roadside scrub lit	[FSCA (1) (TH Atkinson-2007, unpublished)]
48->	United States, Alabama, Hwy 189, 1,5 mi N jct co rd 337; 30/I/2007; Turnbow, R.H.; litter: berlese leaf litter	[RHTC (3) (TH Atkinson-2008, unpublished)]
49->	United States, Alabama, Seale, 3.5 mi NW; 9/II/2002; O'Brien, C.W.; Turnbow, R.H.; litter: berlese mixed hardwood leaf litte	[RHTC (1) (TH Atkinson-2008, unpublished)]
50->	United States, Alabama, Solon Dixon For. Educ. Ctr.; 5/II/2003; O'Brien, C.W.; litter: berlese mixed hardwood litter	[FSCA (2) (TH Atkinson-2007, unpublished)]
51->	United States, Arkansas, Little Rock; 14/X/2001; Baldwin; litter: berlese leaf litter	[TAMU (2) (TH Atkinson-2008, unpublished)]
52->	United States, California, Hermosa	[CAS (1) (Bright, D.E., Stark, R.W. 1973: 69)]
53->	United States, California, Laguna	[Unknown () (Wood, S.L. 1982: 897)]
54->	United States, California, Pasadena	[Unknown () (Wood, S.L. 1982: 897)]
55->	United States, California, Pomona	[MCZ (1) (TH Atkinson-1988, unpublished)]
56->	United States, California, Pomona; 25/IV/1895; Fall, H.C.	[USNM (1: holotype, female) (Wood, S.L. 1982: 897) (Hopkins, A.D. 1915: 19) (TH Atkinson-2015, unpublished) (Wood, S.L. 2007: 517)]
57->	United States, California, Redondo	[Unknown () (Wood, S.L. 1982: 897)]
58->	United States, California, Westwood Hills; 8/V/1925; McClay, A.T.	[MCZ (1) (TH Atkinson-1988, unpublished)]
59->	United States, California, Westwood Hills; Compositae: Encelia farinosa	[EMEC (1) (Bright, D.E., Stark, R.W. 1973: 69)]
60->	United States, District of Columbia, Washington	[Unknown () (Wood, S.L. 1982: 897)]
61->	United States, Florida, 11 mi W Jasper; 24/III/1977; O'Brien, C.W.; O'Brien, L.B.; Marshall, G.B.; litter: berlese	[TAMU (6) (TH Atkinson-1989, unpublished)]
62->	United States, Florida, 8 km NW Starke; 30/VI/1980; Wilkening, A.J.; trap: window trap ethanol	[TAMU (1) (TH Atkinson-1989, unpublished)]
63->	United States, Florida, 8 mi S Marianna; 28/II/1977; O'Brien, C.W.; O'Brien, L.B.; Marshall, G.B.; litter: berlese	[TAMU (4) (TH Atkinson-1989, unpublished)]
64->	United States, Florida, Gainesville	[Unknown () (Wood, S.L. 1982: 897)]
65->	United States, Florida, Homestead	[Unknown () (Wood, S.L. 1982: 897)]
66->	United States, Florida, Hwy 135 at Ochlockonee Riv.; 13/I/2000; O'Brien, C.W.; Marshall, G.B.; litter: berlese hardwood litter	[CAS (1) (TH Atkinson-2007, unpublished)]
67->	United States, Florida, Key Largo	[Unknown () (Wood, S.L. 1982: 897)]
68->	United States, Florida, Key Vaca	[Unknown () (Wood, S.L. 1982: 897)]
69->	United States, Florida, Key West	[Unknown () (Wood, S.L. 1982: 897)]
70->	United States, Florida, Matecumbe Key	[Unknown () (Wood, S.L. 1982: 897)]
71->	United States, Florida, Ocala Natl For 14 mi E Lynne; 25/I/1974; O'Brien, C.W.; litter: berlese	[MCZ (3) (TH Atkinson-1988, unpublished)]
72->	United States, Florida, Perrine	[Unknown () (Wood, S.L. 1982: 897)]
73->	United States, Florida, Plantation Key	[Unknown () (Wood, S.L. 1982: 897)]
74->	United States, Florida, Torreya St. Pk.; 5/V/2007; Turnbow, R.H.	[RHTC (1) (TH Atkinson-2008, unpublished)]
75->	United States, Florida, Windsor; 1/II/1989; Phillips, T.W.; trap: Lindgren Funnel	[TAMU (2) (TH Atkinson-1989, unpublished)]
76->	United States, Florida, Woodville, Natural Wells Rd.; 26/III/2004; Haseeb, M.; Marshall, G.B.; litter: berlese mixed pine hardwood leaf	[FSCA (1) (TH Atkinson-2007, unpublished)]
77->	United States, Georgia, Athens; 15/III/1982; Monkman, K.; Moraceae: Ficus carica	[UGCA (4) (TH Atkinson-1988, unpublished)]
78->	United States, Georgia, Athens, 5 mi W; 30/V/1978; Smith, C.L.; trap: pitfall	[UGCA (1) (TH Atkinson-1988, unpublished)]
79->	United States, Georgia, Dubois, 1.8 mi ESE; 27/XII/2003; Turnbow, R.H.; litter: berlese pine - wiregrass litter	[RHTC (1) (TH Atkinson-2008, unpublished)]
80->	United States, Georgia, Geneva; 9/II/2002; O'Brien, C.W.; Turnbow, R.H.; litter: berlese beech litter	[RHTC (2) (TH Atkinson-2008, unpublished)]
81->	United States, Georgia, Greston, 1.1 mi NE; 11/I/2004; Turnbow, R.H.; litter: berlese leaf litter	[RHTC (2) (TH Atkinson-2008, unpublished)]
82->	United States, Georgia, Hwy 21, 15.9 mi S jct. hwy 301; 22/I/2004; O'Brien, C.W.; Turnbow, R.H.; litter: berlese leaf litter	[RHTC (2) (TH Atkinson-2008, unpublished)]
83->	United States, Georgia, Ichauway Ecol. Res. Ctr.; 22/I/2004; O'Brien, C.W.; Turnbow, R.H.; litter: berlese leaf litter	[RHTC (3) (TH Atkinson-2008, unpublished)]
84->	United States, Georgia, Valdosta, 2.8 mi N; 5/I/2005; O'Brien, C.W.; Haseeb, M.; litter: berlese sifted pine oak litter	[FSCA (1: paratype(s), female) (Johnson, A.J.; Atkinson, T.H.; Hulcr, J. 2016: 421)]
85->	United States, Georgia, Whitehall Forest; 17/V/1978; Smith, C.L.; trap: pitfall	[UGCA (2) (TH Atkinson-1988, unpublished)]
86->	United States, Georgia, Whitehall Forest; 2/V/1979; Smith, C.L.; trap: pitfall	[UGCA (1) (TH Atkinson-1988, unpublished)]
87->	United States, Illinois, Alexander Co.; 1989; Weber, B.C.; trap: window trap	[unknown (1) (Weber, B.C., MacPherson, J.E. 1991: 55)]
88->	United States, Indiana, Clark Co.	[USNM (1: holotype, female) (Wood, S.L.; Bright, D.E. 1992: 912) (TH Atkinson-2018, unpublished) (Wood, S.L. 2007: 517)]
89->	United States, Kansas, Lawrence	[Unknown () (Wood, S.L. 1982: 897)]
90->	United States, Kansas, Wellington	[Unknown () (Wood, S.L. 1982: 897)]
91->	United States, Kentucky, Fulton	[Unknown () (Wood, S.L. 1982: 897)]
92->	United States, Louisiana, Baton Rouge; 11/III/1982; Riley, E.G.	[EGRC (1) (TH Atkinson-1987, unpublished)]
93->	United States, Louisiana, Baton Rouge; 23/III/1982; Riley, E.G.	[EGRC (1) (TH Atkinson-1987, unpublished)]
94->	United States, Louisiana, Baton Rouge; 7/V/1982; Riley, E.G.	[EGRC (1) (TH Atkinson-1987, unpublished)]
95->	United States, Louisiana, Chestnut, 1.8 mi NNW; 24/II/2004; O'Brien, C.W.; Turnbow, R.H.; litter: berlese beech, pine, grass litter	[RHTC (1) (TH Atkinson-2008, unpublished)]
96->	United States, Louisiana, Timbalier Isl; 18/V/1983; Westphal; Wilkes	[LSAM (5) (TH Atkinson-1988, unpublished)]
97->	United States, Maryland, College Pk.	[Unknown () (Wood, S.L. 1982: 897)]
98->	United States, Mississippi, Adaton, 3 mi W; 4/VIII/1997; Schieffer, T.L.; trap: Lindgren Funnel turpentine	[MEM (1) ( : 1)]
99->	United States, Mississippi, Ecru, 1 mi. SE ; 8/XII/1980; Snodgrass, G.L.	[MEM (1) ( : 1)]
100->	United States, Mississippi, State College BWRL; 29/VI/1962; Cross, W.H.	[MEM (1) ( : 1)]
101->	United States, Mississippi, Tishomingo S.P.; 11/IV/1986; MacDonald, S.; litter: berlese litter hardwood forest	[MEM (1) ( : 1)]
102->	United States, Missouri, Charleston	[Unknown () (Wood, S.L. 1982: 897)]
103->	United States, Missouri, Jefferson City	[MCZ (1) (TH Atkinson-1988, unpublished)]
104->	United States, New Jersey, Haddon Heights	[Unknown () (Wood, S.L. 1982: 897)]
105->	United States, North Carolina, McDowell Co.; 1977; Weber, B.C.; trap: window trap	[unknown (1) (Weber, B.C., MacPherson, J.E. 1991: 46)]
106->	United States, Oklahoma, Latimer Co.; 1982; Stephan, K.	[TAMU (26) (Atkinson, T.H.; Riley, E.G. 2013: 7) (Atkinson, T.H.; Riley, E.G. 2013: 7)]
107->	United States, Oklahoma, Latimer Co.; VIII/1992; Stephan, K.	[TAMU (2) (Atkinson, T.H.; Riley, E.G. 2013: 7)]
108->	United States, Oklahoma, Latimer Co.; V/1994; Stephan, K.	[TAMU (1) (Atkinson, T.H.; Riley, E.G. 2013: 7)]
109->	United States, Oklahoma, Latimer Co.; IX/1994; Stephan, K.	[TAMU (1) (Atkinson, T.H.; Riley, E.G. 2013: 7)]
110->	United States, Oklahoma, Red Oak; 15/IX/1978; Stephan, K.	[TAMU (1) (Atkinson, T.H.; Riley, E.G. 2013: 7) (Atkinson, T.H.; Riley, E.G. 2013: 7)]
111->	United States, Oklahoma, Red Oak; IX/1978; Stephan, K.	[TAMU (1) (Atkinson, T.H.; Riley, E.G. 2013: 7) (Atkinson, T.H.; Riley, E.G. 2013: 7)]
112->	United States, South Carolina, Bamberg, 13.4 mi SW; 7/I/2005; O'Brien, C.W.; Turnbow, R.H.; litter: berlese leaf litter	[RHTC (1) (TH Atkinson-2008, unpublished)]
113->	United States, South Carolina, Charleston	[Unknown () (Wood, S.L. 1982: 897)]
114->	United States, South Carolina, Florence Airport; 7/I/2005; O'Brien, C.W.; Turnbow, R.H.; litter: berlese leaf litter	[RHTC (1) (TH Atkinson-2008, unpublished)]
115->	United States, South Carolina, Frogmore	[Unknown () (Wood, S.L. 1982: 897)]
116->	United States, South Carolina, Isle of Palms	[Unknown () (Wood, S.L. 1982: 897)]
117->	United States, South Carolina, Pawley's Beach (Island)	[Unknown () (Wood, S.L. 1982: 897)]
118->	United States, South Carolina, Sand Hills St. For.; 8/I/2005; O'Brien, C.W.; Turnbow, R.H.; litter: berlese leaf litter	[RHTC (1) (TH Atkinson-2008, unpublished)]
119->	United States, South Carolina, Savannah River Site; 9/VIII/2011; Coyle, D.R.; trap: Lindgren Funnel	[UGCA (2) (TH Atkinson-2011, unpublished)]
120->	United States, Tennessee, Chapin Sanctuary, East Ridge	[Unknown () (Wood, S.L. 1982: 897)]
121->	United States, Texas, Alamo	[Unknown () (Wood, S.L. 1982: 897)]
122->	United States, Texas, Altair, 1 mi S; 1/II/1989; Anderson, R.; litter: berlese oak litter	[TAMU (1) (TH Atkinson-2008, unpublished)]
123->	United States, Texas, Angelina NF, 5.5 mi SE Zavala; 17/V/1996; Clarke; Menard; Riley, E.G.; trap: pitfall	[TAMU (1) (TH Atkinson-2006, unpublished)]
124->	United States, Texas, Austin, Guerrero Park; 1/X/2014; Atkinson, T.H.; Apocynaceae: Funastrum clausum	[UTIC (2) (TH Atkinson-2015, unpublished)]
125->	United States, Texas, Big Bend Natl. Pk.	[Unknown () (Wood, S.L. 1982: 897)]
126->	United States, Texas, Boca Chica	[Unknown () (Wood, S.L. 1982: 897)]
127->	United States, Texas, College Station; 3/IV/1932; Reinhard, H.J.	[TAMU (3) (SL Wood-1953, unpublished) (TH Atkinson-1987, unpublished)]
128->	United States, Texas, College Station; 17/III/1932; Reinhard, H.J.	[TAMU (4) (SL Wood-1953, unpublished) (TH Atkinson-1987, unpublished)]
129->	United States, Texas, College Station; 8/V/1932; Reinhard, H.J.	[TAMU (1) (SL Wood-1953, unpublished) (TH Atkinson-1987, unpublished)]
130->	United States, Texas, College Station, Lick Creek Pk.; 3/VII/2004; Dean, D.A.; trap: pitfall	[TAMU (1) (TH Atkinson-2012, unpublished)]
131->	United States, Texas, Dallas; Poaceae: Triticum sp.	[USNM (1: holotype, female) (Wood, S.L. 1982: 897) (Hopkins, A.D. 1915: 19) (Wood, S.L.; Bright, D.E. 1992: 912) (TH Atkinson-2018, unpublished) (Wood, S.L. 2007: 517)]
132->	United States, Texas, Dripping Springs, 6 mi NW; 31/III/2006; Riley, E.G.; trap: flight intercept	[TAMU (3) (TH Atkinson-2006, unpublished)]
133->	United States, Texas, Dripping Springs, 6 mi NW; 24/II/2006; Riley, E.G.; trap: flight intercept	[TAMU (1) (TH Atkinson-2006, unpublished)]
134->	United States, Texas, Dripping Springs, 6 mi NW; 3/VI/2006; Riley, E.G.; trap: flight intercept	[TAMU (3) (TH Atkinson-2006, unpublished)]
135->	United States, Texas, Elkhart, 10 mi SW; 23/II/2004; O'Brien, C.W.; Turnbow, R.H.; litter: berlese leaf litter	[RHTC (5) (TH Atkinson-2008, unpublished)]
136->	United States, Texas, Hemphill, 8.8 mi NE; 16/III/2008; Riley, E.G.; trap: flight intercept	[EGRC (1) (TH Atkinson-2008, unpublished)]
137->	United States, Texas, Kerrville	[Unknown () (Wood, S.L. 1982: 897)]
138->	United States, Texas, New Waverly, 12.2 mi W; 3/VII/1975; Ashe, J.S.; litter: berlese pine litter	[TAMU (1) (TH Atkinson-2008, unpublished)]
139->	United States, Texas, Pharr	[Unknown () (Wood, S.L. 1982: 897)]
140->	United States, Texas, Port Arthur	[Unknown () (Wood, S.L. 1982: 897)]
141->	United States, Texas, Roy Larsen Sandyland Sanct.; 21/II/2004; O'Brien, C.W.; Turnbow, R.H.; litter: berlese longleaf pine, oak litter	[RHTC (1) (TH Atkinson-2008, unpublished)]
142->	United States, Texas, San Antonio; 11/IV/1930; Reinhard, H.J.; Euphorbiaceae: Croton sp.	[TAMU (1) (SL Wood-1953, unpublished)]
143->	United States, Texas, San Jacinto Co.; 27/II/1982; Young, J.K.; litter: berlese	[TAMU (1) (TH Atkinson-1988, unpublished)]
144->	United States, Virginia, Lynchburg	[Unknown () (Wood, S.L. 1982: 897)]
145->	United States, Virginia, Spotsylvania Co.	[Unknown () (Wood, S.L. 1982: 897)]
South America
146->	Brazil, Bahia, Vitória da Conquista; 2/III/2011; Oliveira, L.C.; Moraceae: Artocarpus heterophyllus	[MEFEIS (29) (TH Atkinson-2014, unpublished)] [UTIC (8) (TH Atkinson-2014, unpublished)]
147->	Brazil, Mato Grosso do Sul, Selviria; 2010; Silva, J.C.C.; trap: window trap unbaited	[MEFEIS (1) (TH Atkinson-2014, unpublished)]
148->	Brazil, Paraiba, Patos; 2011; Guedes, R.S.; trap: flight intercept ethanol	[MEFEIS (2) (TH Atkinson-2014, unpublished)]
149->	Brazil, São Paulo, Pinhal	[MZSP (1: holotype, female) (Wood, S.L.; Bright, D.E. 1992: 912) (Wood, S.L. 2007: 517)]
150->	Ecuador, Galápagos, Isla Española	[MSUC (0) (Bright, D.E., Skidmore, R.E. 2002: 141)]
151->	Ecuador, Galápagos, Isla Fernandina	[MSUC (0) (Bright, D.E., Skidmore, R.E. 2002: 141)]
152->	Ecuador, Galápagos, Isla Floreana	[MSUC (0) (Bright, D.E., Skidmore, R.E. 2002: 141)]
153->	Ecuador, Galápagos, Isla San Cristobal	[MSUC (0) (Bright, D.E., Skidmore, R.E. 2002: 141)]
154->	Ecuador, Galápagos, Isla Sta. Cruz	[MSUC (0) (Bright, D.E., Skidmore, R.E. 2002: 141)]
UNK
155->	, no state / prov. data; Liliaceae: Aloe vera	[unknown (0) (Wood, S.L. 1982: 897)]
156->	, no state / prov. data; Leguminosae: Bauhinia alba	[unknown (0) (Wood, S.L. 1982: 897)]
157->	, no state / prov. data; Compositae: Bidens pilosa	[unknown (0) (Wood, S.L. 1982: 897)]
158->	, no state / prov. data; Leguminosae: Cajanus cajon	[unknown (0) (Wood, S.L. 1982: 897)]
159->	, no state / prov. data; Scrophulariaceae: Capraria biflora	[unknown (0) (Wood, S.L. 1982: 897)]
160->	, no state / prov. data; Leguminosae: Galactia spiciformis	[unknown (0) (Wood, S.L. 1982: 897)]
161->	, no state / prov. data; Convolvulaceae: Ipomoea cathartica	[unknown (0) (Wood, S.L. 1982: 897)]
162->	, no state / prov. data; Convolvulaceae: Ipomoea littoralis	[unknown (0) (Wood, S.L. 1982: 897)]
163->	, no state / prov. data; Compositae: Iva imbricata	[unknown (0) (Wood, S.L. 1982: 897)]
164->	, no state / prov. data; Malvaceae: Malvastrum sp.	[unknown (0) (Wood, S.L. 1982: 897)]
165->	, no state / prov. data; Anacardiaceae: Mangifera indica	[unknown (0) (Wood, S.L. 1982: 897)]
166->	, no state / prov. data; Poaceae: Paspalum vaginatum	[unknown (0) (Wood, S.L. 1982: 897)]
167->	, no state / prov. data; Combretaceae: Quisqualis indica	[unknown (0) (Wood, S.L. 1982: 897)]
168->	, no state / prov. data; Salicaceae: Salix babylonica	[unknown (0) (Wood, S.L. 1982: 897)]
169->	, no state / prov. data; Poaceae: Uniola paniculata	[unknown (0) (Wood, S.L. 1982: 897)]
170->	, no state / prov. data; Poaceae: Zea mays	[unknown (0) (Wood, S.L. 1982: 897)]
171->	, no state / prov. data; Verbenaceae: Verbena sp.	[unknown (0) (Wood, S.L. 1982: 897)]