Bark and Ambrosia Beetles of North and Central America

SYNONYMY

• Scolytus mali (Bechstein 1805) Vollstandige Natur. Schadl. Forstinsekt. 3: 882
• Bostrichus mali Bechstein 1805. Vollstandige Natur. Schadl. Forstinsekt. 3: 822
• Eccoptogaster castaneus Ratzeburg 1837. Die Forst-insekten: 187
• Eccoptogaster pruni Ratzeburg 1837. Die Forst-insekten: 187
• Eccoptogaster pyri Ratzeburg 1837. Die Forst-insekten: 186
• Scolytus sulcatus LeConte 1868. Trans. Amer. Ent. Soc. 2: 167
• Scolytus nitidulus Chapuis 1869. Synopsis des Scolytides: 59
• Scolytus mali strigilatus Reitter 1913. Wiener Ent. Zeit. 32: 19
• Scolytus bicallosus Eggers 1933. Ent. Nachrichtenbl. 7: 75

DISTRIBUTION.

Numbers in parentheses after each geographic unit are the number of distinct collection events in the database for that unit. For exotic species generally only countries are listed for localities outside the New World. For further information on published sources of distribution, check the REFERENCES section.

• (4)
  (4): no state / prov. data (4);
• Africa (3)
  Algeria (1): no state / prov. data (1); Egypt (1): no state / prov. data (1); Morocco (1): no state / prov. data (1);
• Eurasia (36)
  (1): no state / prov. data (1); Armenia (1): no state / prov. data (1); Austria (1): no state / prov. data (1); Azerbaijan (1): no state / prov. data (1); Belgium (1): no state / prov. data (1); Bulgaria (1): no state / prov. data (1); Czech Republic (1): no state / prov. data (1); Czechoslovakia (1): no state / prov. data (1); Denmark (1): no state / prov. data (1); England (1): no state / prov. data (1); Finland (1): no state / prov. data (1); France (1): no state / prov. data (1); Georgia (1): no state / prov. data (1); Germany (4): no state / prov. data (4); Greece (1): no state / prov. data (1); Hungary (1): no state / prov. data (1); Ireland (1): no state / prov. data (1); Italy (1): no state / prov. data (1); Luxemburg (1): no state / prov. data (1); Netherlands (1): no state / prov. data (1); Norway (1): no state / prov. data (1); Poland (1): no state / prov. data (1); Romania (1): no state / prov. data (1); Russia (5): no state / prov. data (5); Spain (1): no state / prov. data (1); Sweden (1): no state / prov. data (1); Switzerland (1): no state / prov. data (1); Turkey (1): no state / prov. data (1); Yugoslavia (1): no state / prov. data (1);
• North America (306)
  Canada (8): Ontario (6), Quebec (2); United States (298): no state / prov. data (3), Alabama (6), Arkansas (4), Connecticut (1), Delaware (3), District of Columbia (1), Florida (2), Georgia (17), Illinois (14), Indiana (27), Iowa (2), Kansas (5), Kentucky (4), Louisiana (7), Maine (1), Maryland (14), Michigan (20), Mississippi (12), Missouri (16), Nebraska (2), New Jersey (9), New York (12), North Carolina (12), Ohio (13), Oklahoma (14), Pennsylvania (25), South Carolina (8), Tennessee (5), Texas (29), Vermont (1), Virginia (5), West Virginia (3), Wisconsin (1);
• UNK (2)
  (2): no state / prov. data (2);

HOSTS

Numbers in parentheses after each host family, genus, or collection method are the number of distinct collection events in the database for that host or method.

• No host information (121);
• Betulaceae (2): Betula (1): nigra (1); Ostrya (1): virginiana (1);
• Cannabaceae (2): Celtis (2): lindheimeri (1), sp. (1);
• Fagaceae (4): Castanea (1): dentata (1); Fagus (1): grandifolia (1); Quercus (2): rubra (1), velutina (1);
• Juglandaceae (21): Carya (20): glabra (1), illinoensis (2), ovata (7), sp. (10); Juglans (1): nigra (1);
• Leguminosae (1): Cladrastris (1): lutea (1);
• Magnoliaceae (1): Liriodendron (1): tulipifera (1);
• Rosaceae (11): Cotoneaster (1): sp. (1); Crataegus (1): sp. (1); Cydonia (1): sp. (1); Malus (5): pumila (4), sp. (1); Prunus (1): sp. (1); Pyrus (1): sp. (1); Sorbus (1): sp. (1);
• Sapindaceae (2): Acer (2): saccharum (1), sp. (1);
• trap (183): (91): no details (1), alpha pinene + ethanol (90); alpha-scents 4 vane (1): ethanol (1); bottle (2): ethanol (2); bottle trap (2): no details (2); canopy (3): no details (3); flight intercept (2): no details (2); light (4): no details (1), ultraviolet (3); Lindgren Funnel (68): no details (20), alpha pinene + ethanol (9), ethanol (25), exotic ips lure (11), turpentine (2), turpentine + frontalin (1); malaise (1): no details (1); purple prism (2): no details (2); soda bottle (1): ethanol (1); sugar trap (1): no details (1); window trap (4): no details (2), ethanol (2); yellow pan trap (1): no details (1);
• Ulmaceae (3) Ulmus (3): americana (2), sp. (1);

REFERENCES

The following are important recent monographs, catalogs, and supplements to catalogs that refer to this species. The specific page on which the reference is made is shown in pink at the end of the reference. In the case of Wood (1982) and Wood & Bright (1992) clicking on the reference page will link to a digital version of the work in question.

• Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6:1-1356. [426]. (data capture complete)
• Wood, S.L., Bright,D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index. Great Basin Nat. Mem. 13:1-1553 (vol. A, B). [341]

The following are references from which host and distribution data have been input into the database. If one of the above monographs or catalogs also appears in this list, it means that most relevant collection event data have been included.

1. . . .
2. Anonymous, 2010. Michigan State University. SCAN: Symbiota Collections of Artropods Network. http://symbiota4.acis.ufl.edu/scan/portal/collections/misc/collprofiles.php?collid=76: -.
3. Bright, D.E.; Skidmore, R.S. 1997. A Catalog of Scolytidae and Platypodidae (Coleoptera), Supplement 1 (1990-1994). A Catalog of Scolytidae and Platypodidae (Coleoptera), Supplement 1 (1990-1994): -.
4. Cognato, A.I.; Barc, N.; Philip, M.; Mech, R.; Smith, A.D.; Galbraith, E.; Storer, A.J.; Kirkendall, L.R. 2009. The native and introduced bark and ambrosia beetles of Michigan (Curculionidae: Scolytinae). Great Lakes Entom.42(3-4): 101-120.
5. Deyrup, M.A. 1981. Annotated list of Indiana Scolytidae (Coleoptera). Great Lakes Entomol. 14: 1-9.
6. EDRR 2011. USFS Early Detection Rapid Response Database. http://www.fs.fed.us/invasivespecies/earlydetection.shtml/.
7. Petrov, A.V. 2013. New data and synonymy of Scolytus Geoffroy, 1763 bark beetles ( Coleoptera: Curculionidae: Scolytinae( from Russia and adjacent countries. Lesnoy Vetsnik 98(6): 39-47.
8. Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6: 1-1356.
9. Wood, S.L.; Bright, D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index (Volumes A,B). Great Basin Nat. Mem. 13: 1-1553.

Maps automatically open at the center of the plotted points and the scale is set to encompass all map points. Maps can be resized (scale bar at upper left) and the center moved (place cursor over map and drag) to see other parts of the distribution of the species. Clicking on a map marker will pull up collection event data and a literature citation if present. If any errors are found, please refer to the "series code" which is a unique identifier for a database record in any communications). Coordinates have not been entered for all collection records. Localities outside the New World are not plotted, even though they are listed in the distribution summary and in the table of records.

No.	Collection data	Publication
1->	, no state / prov. data; Rosaceae: Prunus sp.	[unknown (0) (Wood, S.L. 1982: 426)]
2->	, no state / prov. data; Rosaceae: Pyrus sp.	[unknown (0) (Wood, S.L. 1982: 426)]
3->	, no state / prov. data; Rosaceae: Cotoneaster sp.	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 341)]
4->	, no state / prov. data; Rosaceae: Cydonia sp.	[unknown (0) (Petrov, A.V. 2013: 43)]
Africa
5->	Algeria, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
6->	Egypt, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
7->	Morocco, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
Eurasia
8->	, no state / prov. data	[TAMU (2) (TH Atkinson-, unpublished)]
9->	Armenia, no state / prov. data, Armenia (country)	[unknown (1) (Petrov, A.V. 2013: 43)]
10->	Austria, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
11->	Azerbaijan, no state / prov. data, Azerbaijan (country)	[unknown (1) (Petrov, A.V. 2013: 43)]
12->	Belgium, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
13->	Bulgaria, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
14->	Czech Republic, no state / prov. data, Czech Republic (country)	[unknown (1) (Petrov, A.V. 2013: 43)]
15->	Czechoslovakia, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
16->	Denmark, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
17->	England, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
18->	Finland, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
19->	France, no state / prov. data, 'France meridionale'	[IRSNB (1: holotype, female) (Wood, S.L.; Bright, D.E. 1992: 341)]
20->	Georgia, no state / prov. data, Georgia (country)	[unknown (1) (Petrov, A.V. 2013: 43)]
21->	Germany, no state / prov. data	[unknown (1: syntype(s)) (Wood, S.L.; Bright, D.E. 1992: 341)]
22->	Germany, no state / prov. data	[ZIFH (1: syntype(s)) (Wood, S.L.; Bright, D.E. 1992: 341)]
23->	Germany, no state / prov. data	[ZIFH (1: syntype(s)) (Wood, S.L.; Bright, D.E. 1992: 341)]
24->	Germany, no state / prov. data, Lothgangen	[ZIFH (1: syntype(s)) (Wood, S.L.; Bright, D.E. 1992: 341)]
25->	Greece, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
26->	Hungary, no state / prov. data, Sagvar	[USNM (1: holotype, male) (Wood, S.L.; Bright, D.E. 1992: 341) (Anderson, W.H.; Anderson, D.M. 1971: 6)]
27->	Ireland, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
28->	Italy, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
29->	Luxemburg, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
30->	Netherlands, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
31->	Norway, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
32->	Poland, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
33->	Romania, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
34->	Russia, no state / prov. data, Eastern Siberia	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341) (Petrov, A.V. 2013: 43)]
35->	Russia, no state / prov. data, Khabarovsk Krai	[unknown (1) (Petrov, A.V. 2013: 43)]
36->	Russia, no state / prov. data, northern Russia	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341) (Petrov, A.V. 2013: 43)]
37->	Russia, no state / prov. data, southern Primorye	[unknown (1) (Petrov, A.V. 2013: 43)]
38->	Russia, no state / prov. data, Transbaikalia	[unknown (1) (Petrov, A.V. 2013: 43)]
39->	Spain, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
40->	Sweden, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
41->	Switzerland, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
42->	Turkey, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
43->	Yugoslavia, no state / prov. data	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
North America
44->	Canada, Ontario, Middle Island; 4/VII/2000; Paiero, S.M.; Marshall, S.A.; Cheung; trap: yellow pan trap	[DEBU (4) (Douglas, H.; Bouchard, P.; Anderson, R.S.; De Tonnancour, P.; Vigneault, R.; Webster, P. 2013: 39)]
45->	Canada, Ontario, Middle Island; 11/VI/2003; Marshall, S.A.	[DEBU (1) (Douglas, H.; Bouchard, P.; Anderson, R.S.; De Tonnancour, P.; Vigneault, R.; Webster, P. 2013: 39)]
46->	Canada, Ontario, Oakville; 9/VI/2008; CFIA; trap: Lindgren Funnel	[CNC (1) (Douglas, H.; Bouchard, P.; Anderson, R.S.; De Tonnancour, P.; Vigneault, R.; Webster, P. 2013: 39)]
47->	Canada, Ontario, Pt. Pelee N.P.; 22/V/2000; Lonsdale, O.	[DEBU (1) (Douglas, H.; Bouchard, P.; Anderson, R.S.; De Tonnancour, P.; Vigneault, R.; Webster, P. 2013: 39)]
48->	Canada, Ontario, Pt. Pelee N.P., The Tip; 17/VI/2003; Carscadden, H.	[DEBU (1) (Douglas, H.; Bouchard, P.; Anderson, R.S.; De Tonnancour, P.; Vigneault, R.; Webster, P. 2013: 39)]
49->	Canada, Ontario, southern Ontario	[unknown () (Wood, S.L. 1982: 426)]
50->	Canada, Quebec, Oka, Parc d'Oka, Deux-Montagnes; 27/IV/1997; Vigneault, R.; Juglandaceae: Carya ovata	[CRVI (1) (Douglas, H.; Bouchard, P.; Anderson, R.S.; De Tonnancour, P.; Vigneault, R.; Webster, P. 2013: 39)]
51->	Canada, Quebec, Quebec (province)	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 341)]
52->	United States, no state / prov. data; Fagaceae: Fagus grandifolia	[unknown (0) (Wood, S.L. 1982: 614)]
53->	United States, no state / prov. data; Fagaceae: Quercus velutina	[unknown (0) (Wood, S.L. 1982: 614)]
54->	United States, no state / prov. data; Ulmaceae: Ulmus americana	[unknown (0) (Wood, S.L. 1982: 614)]
55->	United States, Alabama, Corona Mines; 18/IX/1976; Hawkins, B.A.	[Unknown () (Wood, S.L. 1982: 615)]
56->	United States, Alabama, Highland Lake; 13/VII/2012; King, T.; trap: light	[FSCA (1) (TH Atkinson-2021, unpublished)]
57->	United States, Alabama, Highland Lake; 10/VI/2012; King, T.; trap: window trap	[FSCA (1) (TH Atkinson-2021, unpublished)]
58->	United States, Alabama, Lincoln; 28/III/2008; McReynolds, D.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
59->	United States, Alabama, Lincoln; 24/IV/2008; trap	[UTIC (1) (TH Atkinson-2020, unpublished)]
60->	United States, Alabama, Lincoln; 10/VI/2008; trap: Lindgren Funnel ethanol	[UTIC (1) (TH Atkinson-2020, unpublished)]
61->	United States, Arkansas, Batesville; 3/IV/2009; Northum, J.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
62->	United States, Arkansas, Forrest City; 15/IV/2009; Northum, J.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
63->	United States, Arkansas, Little Rock; 8/V/1997; Baldwin; trap: sugar trap	[TAMU (1) (TH Atkinson-2002, unpublished)]
64->	United States, Arkansas, Little Rock; 15/V/1997; Baldwin; trap: flight intercept	[TAMU (1) (TH Atkinson-2012, unpublished)]
65->	United States, Connecticut, Connecticut (state)	[unknown () (Wood, S.L. 1982: 426)]
66->	United States, Delaware, Elsmere; 24/IV/2009; Gladders, G.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
67->	United States, Delaware, Killens Pond St. Pk.; 1/VII/1998; Valenti, M.A.	[RJRC (1) (Rabaglia, R.J.; Valenti, M.A. 2003: 314)]
68->	United States, Delaware, Smyrna; 27/IV/2009; Gladders, G.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
69->	United States, District of Columbia, DC	[USNM () (Blackman, M.W. 1928: 454) (Wood, S.L. 1982: 614) (Atkinson, T.H. 1989: 327)]
70->	United States, Florida, Suwannee River S.P.; 25/IX/2009; Whilby, L.; trap: purple prism	[FSCA (8) (TH Atkinson-2010, unpublished)] [THAC (2) (TH Atkinson-2010, unpublished)]
71->	United States, Florida, Suwannee River S.P.; 25/IX/2009; Whilby, L.; trap: purple prism	[FSCA (2) (TH Atkinson-2010, unpublished)]
72->	United States, Georgia, Bishop; 16/IV/2014; Barnes, B.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
73->	United States, Georgia, Black Mtn.	[Unknown () (Wood, S.L. 1982: 614)]
74->	United States, Georgia, Black Mtn., 28 m SW Clayton; 31/VII/1964; Cambre, L.; trap: light ultraviolet	[FSCA (3) (CW O'Brien-2003, unpublished) (TH Atkinson-2007, unpublished)]
75->	United States, Georgia, Clarkesville; 8/I/1909	[CNC (1) (Blackman, M.W. 1920: 449) (Wood, S.L. 1982: 614)]
76->	United States, Georgia, Fulton; 5/IV/2011; Price, T.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
77->	United States, Georgia, Hull; 1/IV/2011; Brownell, K.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
78->	United States, Georgia, Jefferson; 8/IV/2011; Price, T.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
79->	United States, Georgia, Savannah; 16/IV/2013; Morgan, R.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
80->	United States, Georgia, Savannah; 22/V/2012; Morgan, R.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
81->	United States, Georgia, Statham; 1/IV/2011; Brownell, K.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
82->	United States, Georgia, Watkinsville; 21/IV/2010; Brownell, K.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
83->	United States, Georgia, Watkinsville; 8/VI/2012; Adair, J.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
84->	United States, Georgia, Whitehall Forest; 30/V/1975; Turnbow, R.H.; Juglandaceae: Carya sp.	[TAMU (3) (Atkinson, T.H. 1989: 327)] [UGCA (5) (Atkinson, T.H. 1989: 327)]
85->	United States, Georgia, Winder; 22/V/1975; Turnbow, R.H.	[TAMU (1) (Atkinson, T.H. 1989: 327)] [UGCA (1) (Atkinson, T.H. 1989: 327)]
86->	United States, Georgia, Winterville; 21/IV/2010; Brownell, K.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
87->	United States, Georgia, Winterville; 16/IV/2014; Barnes, B.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
88->	United States, Georgia, Winterville; 18/IV/2013; Barnes, B.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
89->	United States, Illinois; 2009; Helm, C.; Molano-Flores, B.; trap: Lindgren Funnel	[INHS (1) (Helm, C.; Molano-Flores, B. 2015: 143)]
90->	United States, Illinois; 2009; Helm, C.; Molano-Flores, B.; trap: Lindgren Funnel	[INHS (1) (Helm, C.; Molano-Flores, B. 2015: 143)]
91->	United States, Illinois; 2009; Helm, C.; Molano-Flores, B.; trap: Lindgren Funnel	[INHS (1) (Helm, C.; Molano-Flores, B. 2015: 143)]
92->	United States, Illinois; 2009; Helm, C.; Molano-Flores, B.; trap: Lindgren Funnel	[INHS (5) (Helm, C.; Molano-Flores, B. 2015: 143)]
93->	United States, Illinois; 2009; Helm, C.; Molano-Flores, B.; trap: Lindgren Funnel	[INHS (2) (Helm, C.; Molano-Flores, B. 2015: 143)]
94->	United States, Illinois; 2009; Helm, C.; Molano-Flores, B.; trap: Lindgren Funnel	[INHS (1) (Helm, C.; Molano-Flores, B. 2015: 143)]
95->	United States, Illinois, Alexander Co.; 14/IX/1979; Weber, B.C.	[USNM (1) (Atkinson, T.H.; Rabaglia,R.J.; Peck,S.J.; Foltz,J.L. 1991: 157)]
96->	United States, Illinois, Effingham; 3/VI/2014; Miller, F.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
97->	United States, Illinois, Homewood; 1/V/2010; Doerr, B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
98->	United States, Illinois, Lemont; 8/V/2009; Miller, F.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
99->	United States, Illinois, Marshall; 13/VI/2014; Miller, F.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
100->	United States, Illinois, Pine Hills Field Station; 15/V/1967	[Unknown () (Wood, S.L. 1982: 615)]
101->	United States, Illinois, Rosedale	[unknown () (Atkinson, T.H. 1989: 327)]
102->	United States, Illinois, West Frankfort; 6/VI/2014; Miller, F.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
103->	United States, Indiana; Juglandaceae: Carya glabra	[unknown (1) (Deyrup, M.A. 1981: 5)]
104->	United States, Indiana; Betulaceae: Ostrya virginiana	[unknown (1) (Deyrup, M.A. 1981: 5)]
105->	United States, Indiana; Sapindaceae: Acer saccharum	[unknown (1) (Deyrup, M.A. 1981: 5)]
106->	United States, Indiana; Fagaceae: Quercus rubra	[unknown (1) (Deyrup, M.A. 1981: 5)]
107->	United States, Indiana; Ulmaceae: Ulmus sp.	[unknown (1) (Deyrup, M.A. 1981: 5)]
108->	United States, Indiana; Leguminosae: Cladrastris lutea	[unknown (1) (Deyrup, M.A. 1981: 5)]
109->	United States, Indiana; Rosaceae: Malus pumila	[unknown (1) (Deyrup, M.A. 1981: 5)]
110->	United States, Indiana; Ulmaceae: Ulmus americana	[unknown (1) (Deyrup, M.A. 1981: 5)]
111->	United States, Indiana, Anderson; 27/V/2008; Stephens, J.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
112->	United States, Indiana, Brown Co. St. Pk.; 5/VI/1981; Deyrup, M.A.; Deyrup, N.; Juglandaceae: Carya ovata	[TAMU (2) (TH Atkinson-2009, unpublished)]
113->	United States, Indiana, Clay Co.	[Unknown () (Deyrup, M.A. 1981: 5)]
114->	United States, Indiana, Columbia City; 13/V/2008; Kruse, R.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
115->	United States, Indiana, Evansville; 20/VIII/2014; Rust, A.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
116->	United States, Indiana, Green Hill; 29/VIII/1981; Deyrup, M.A.; Juglandaceae: Carya ovata	[TAMU (2) (Deyrup, M.A. 1981: 5) (Deyrup, M.A. 1981: 5)]
117->	United States, Indiana, Indianapolis; 26/V/1967; White, C.E.	[FSCA (1) (Atkinson, T.H. 1989: 327)]
118->	United States, Indiana, Jasper-Pulaski WMA; 24/V/1998; Grundel, R.; trap: Lindgren Funnel	[FSCA (1) (TH Atkinson-2021, unpublished)]
119->	United States, Indiana, Marion Co.	[Unknown () (Deyrup, M.A. 1981: 5)]
120->	United States, Indiana, Poland; 1/IV/1957; White, C.E.; Juglandaceae: Carya sp.	[TAMU (2) (Atkinson, T.H. 1989: 327)] [FSCA (1) (Atkinson, T.H. 1989: 327)]
121->	United States, Indiana, Pulaski Co.	[unknown (1) (Deyrup, M.A. 1981: 5)]
122->	United States, Indiana, Shakamak ST. Pk.; Deyrup, M.A.; Juglandaceae: Carya ovata	[ABS (1) (Deyrup, M.A.; Atkinson, T.H. 1987: 67)]
123->	United States, Indiana, Steuben Co.	[unknown (1) (Deyrup, M.A. 1981: 5)]
124->	United States, Indiana, Tippecanoe Co	[unknown (1) (Deyrup, M.A. 1981: 5)]
125->	United States, Indiana, Tippecanoe Co.	[MCZ (1) (TH Atkinson-1988, unpublished)]
126->	United States, Indiana, Tippecanoe Co.; 1/VI/1983; Deyrup, M.A.	[FSCA (1) (Deyrup, M.A. 1981: 5)]
127->	United States, Indiana, Turkey Run St. Pk.; Deyrup, M.A.; Juglandaceae: Carya ovata	[ABS (1) (Deyrup, M.A. 1981: 5) (Deyrup, M.A.; Atkinson, T.H. 1987: 67)]
128->	United States, Indiana, W. Lafayette; 26/III/1982; Deyrup, M.A.; Deyrup, N.; Juglandaceae: Carya ovata	[ABS (1) (Deyrup, M.A.; Atkinson, T.H. 1987: 67) (Deyrup, M.A. 1981: 5)] [TAMU (5) (TH Atkinson-2009, unpublished)]
129->	United States, Indiana, W. Lafayette; 19/VI/1979; Deyrup, M.A.; Deyrup, N.	[TAMU (1) (MA Deyrup-1989, unpublished) (TH Atkinson-1989, unpublished)]
130->	United States, Iowa, Jester Park; 28/IV/1985; Wappes, J.E.	[JEWC (1) (Atkinson, T.H. 1989: 327)]
131->	United States, Iowa, Saylorville; 30/VI/1983; Schieferstein, R.H.	[JEWC (1) (Atkinson, T.H. 1989: 327)]
132->	United States, Kansas, Galena; 12/V/2009; Salsbury; trap alpha pinene + ethanol	[unknown (15) (EDRR 2011: 0)]
133->	United States, Kansas, Galena, 2 mi S; 4/V/2010; Salsbury, G.; trap: canopy	[SEMC (2) (TH Atkinson-2020, unpublished)]
134->	United States, Kansas, Jingo, 3 mi E; 4/V/2010; Salsbury, G.; trap: canopy	[SEMC (1) (TH Atkinson-2020, unpublished)]
135->	United States, Kansas, Ottawa	[Unknown () (Wood, S.L. 1982: 614)]
136->	United States, Kansas, Uniontown, 1 mi S; 4/V/2010; Salsbury, G.; trap: canopy	[SEMC (1) (TH Atkinson-2020, unpublished)]
137->	United States, Kentucky, Calloway Co.; 2016; trap: bottle ethanol	[unknown (1) (Viloria, Z.; Villanueva, R.T.; Bessin, R.; O'Neal, P.; Ranger, C.M.; Dunwell, W. 2021: 378)]
138->	United States, Kentucky, Georgetown; 29/V/2012; Thompson, J.M.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
139->	United States, Kentucky, Humble	[Unknown () (Wood, S.L. 1982: 614)]
140->	United States, Kentucky, Todd Co.; 2016; trap: bottle ethanol	[unknown (1) (Viloria, Z.; Villanueva, R.T.; Bessin, R.; O'Neal, P.; Ranger, C.M.; Dunwell, W. 2021: 378)]
141->	United States, Louisiana, Alexandria; 1/III/2010; Haley, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
142->	United States, Louisiana, Algiers; 18/IV/1945; Rau; Juglandaceae: Carya illinoensis	[USNM (1: paratype(s), female) (Wood, S.L. 1982: 614) (Atkinson, T.H. 1989: 335) (Wood, S.L. 1974: 17)]
143->	United States, Louisiana, Baton Rouge; 10/VIII/1986; Riley, E.G.; Juglandaceae: Carya sp.	[EGRC (6) (TH Atkinson-1987, unpublished)] [TAMU (2) (TH Atkinson-1987, unpublished)] [LSAM (1) (TH Atkinson-2013, unpublished)]
144->	United States, Louisiana, Baton Rouge; 10/VII/1986; Riley, E.G.	[LSAM (1) (TH Atkinson-1988, unpublished)] [TAMU (1) (TH Atkinson-1988, unpublished)]
145->	United States, Louisiana, Shreveport; 1/III/2010; Haley, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
146->	United States, Louisiana, St. Joseph; 20/III/2012; Haley, T.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
147->	United States, Louisiana, St. Joseph; 20/III/2012; Haley, T.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
148->	United States, Maine, Maine (state)	[unknown (0) (Bright, D.E.; Skidmore, R.S. 1997: 78)]
149->	United States, Maryland, Frederick; 28/III/2011; Lupp, T.; Firme, C.A.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
150->	United States, Maryland, Halethorpe; 25/III/2011; CK; LRT; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
151->	United States, Maryland, Ijamsville; 28/III/2011; Lupp, T.; Firme, C.A.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
152->	United States, Maryland, Jackson's Isl	[USNM () (Blackman, M.W. 1928: 454) (Wood, S.L. 1982: 614) (Atkinson, T.H. 1989: 327)]
153->	United States, Maryland, Maryland (state)	[unknown () (Wood, S.L. 1982: 426)]
154->	United States, Maryland, North Potomac; 7/V/2011; Wappes, J.E.	[FSCA (1) (TH Atkinson-2021, unpublished)]
155->	United States, Maryland, Plummer's Island	[USNM () (Blackman, M.W. 1928: 454) (Atkinson, T.H. 1989: 327)]
156->	United States, Maryland, Plummer's Island; Hopkins, A.D.; Cannabaceae: Celtis sp.	[USNM () (Wood, S.L. 1982: 615) (Atkinson, T.H. 1989: 327)]
157->	United States, Maryland, Plummer's Island; Hopkins, A.D.; Juglandaceae: Carya sp.	[USNM () (Wood, S.L. 1982: 615) (Atkinson, T.H. 1989: 327)]
158->	United States, Maryland, St. Michaels; 7/IV/2011; Tilley, S.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
159->	United States, Maryland, Upper Marlboro; 31/III/2011; Tatman, R.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
160->	United States, Maryland, Wye; 12/V/1995; Rabaglia, R.J.; trap: Lindgren Funnel	[TAMU (2) (RJ Rabaglia-1995, unpublished) (TH Atkinson-2007, unpublished)]
161->	United States, Maryland, Wye; 19/V/1995; Rabaglia, R.J.; trap: Lindgren Funnel	[TAMU (1) (RJ Rabaglia-1995, unpublished) (TH Atkinson-2007, unpublished)]
162->	United States, Maryland, Wye; 26/V/1994; Rabaglia, R.J.; trap: Lindgren Funnel	[TAMU (1) (RJ Rabaglia-1995, unpublished) (TH Atkinson-2007, unpublished)]
163->	United States, Michigan, Detroit; 1874; Hubbard, H.G.; Schwarz, E.A.	[MCZ (1: holotype, female) (Blackman, M.W. 1920: 449) (Wood, S.L. 1982: 614) (Atkinson, T.H. 1989: 327)]
164->	United States, Michigan, E. State Rd., 2 mi N of Lansing; 6/II/2007; Parsons, G.L.	[MSUC (1) (Cognato, A.I.; Barc, N.; Philip, M.; Mech, R.; Smith, A.D.; Galbraith, E.; Storer, A.J.; Kirkendall, L.R. 2009: 1) (Anonymous, 2010: 0)]
165->	United States, Michigan, E.S. George Res.; 17/IX/1978; Kirkendall, L.R.; Rosaceae: Malus pumila	[unknown (1) (Cognato, A.I.; Barc, N.; Philip, M.; Mech, R.; Smith, A.D.; Galbraith, E.; Storer, A.J.; Kirkendall, L.R. 2009: 1)]
166->	United States, Michigan, Farmington Hills; 7/II/2004; Sullivan, B.T.	[MSUC (1) (Cognato, A.I.; Barc, N.; Philip, M.; Mech, R.; Smith, A.D.; Galbraith, E.; Storer, A.J.; Kirkendall, L.R. 2009: 1) (Anonymous, 2010: 0)]
167->	United States, Michigan, Fennville area; 30/VI/2003; McGhee, P.; Rosaceae: Malus pumila	[MSUC (1) (Cognato, A.I.; Barc, N.; Philip, M.; Mech, R.; Smith, A.D.; Galbraith, E.; Storer, A.J.; Kirkendall, L.R. 2009: 1) (Anonymous, 2010: 0)]
168->	United States, Michigan, Forest Hill Natural Area; 17/VI/2008	[MSUC (1) (Anonymous, 2010: 0)]
169->	United States, Michigan, Gerrish; 5/V/2008; Radtke; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
170->	United States, Michigan, Grosse Ile, Pke Lane; 23/V/2007	[MSUC (1) (Anonymous, 2010: 0)]
171->	United States, Michigan, Grosse lle; 23/V/2007; Mech, R.; trap alpha pinene + ethanol	[unknown (3) (EDRR 2011: 0)]
172->	United States, Michigan, Grosse lle; 23/V/2007; Mech, R.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
173->	United States, Michigan, Howell; 13/V/2011; Mech, R.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
174->	United States, Michigan, Jackson Co.; 13/V/1951; Dreisbach	[MSUC (1) (Anonymous, 2010: 0)]
175->	United States, Michigan, Okemos; 28/V/1970; Wellso, S.G.	[MSUC (1) (Anonymous, 2010: 0)]
176->	United States, Michigan, Pinckney; 12/VII/1981; Deyrup, M.A.; Juglandaceae: Carya ovata	[ABS (1) (Atkinson, T.H. 1989: 335)] [UTIC (4) (Bright, D.E.; Skidmore, R.S. 1997: 96) (Atkinson, T.H. 1989: 335)] [ABS (2) (Atkinson, T.H. 1989: 335)] [ABS (2) (Atkinson, T.H. 1989: 335)] [TAMU (6) (Atkinson, T.H. 1989: 335)]
177->	United States, Michigan, Pine; 5/V/2008; Mech, R.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
178->	United States, Michigan, Richfield; 5/V/2008; Mech, R.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
179->	United States, Michigan, Richfield County Park; 16/VI/2008; Mech, R.	[MSUC (1) (Cognato, A.I.; Barc, N.; Philip, M.; Mech, R.; Smith, A.D.; Galbraith, E.; Storer, A.J.; Kirkendall, L.R. 2009: 1) (Anonymous, 2010: 0)]
180->	United States, Michigan, St. Charles; 25/VI/1969; Truchan, J.G.	[MSUC (1) (Cognato, A.I.; Barc, N.; Philip, M.; Mech, R.; Smith, A.D.; Galbraith, E.; Storer, A.J.; Kirkendall, L.R. 2009: 1) (Anonymous, 2010: 0)]
181->	United States, Michigan, St. Charles; 7/VIII/1968; Truchan, J.G.	[MSUC (1) (Cognato, A.I.; Barc, N.; Philip, M.; Mech, R.; Smith, A.D.; Galbraith, E.; Storer, A.J.; Kirkendall, L.R. 2009: 1) (Anonymous, 2010: 0)]
182->	United States, Michigan, Williamston; 1/V/2011; Cognato, A.I.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
183->	United States, Mississippi, Adaton, 3 mi W; 24/IV/1998; Schieffer, T.L.; trap: Lindgren Funnel	[MEM (2) (TH Atkinson-2009, unpublished)]
184->	United States, Mississippi, Adaton, 3 mi W; 10/X/1997; Schieffer, T.L.; trap: Lindgren Funnel turpentine	[MEM (1) (TH Atkinson-2009, unpublished)]
185->	United States, Mississippi, Adaton, 3 mi W; 13/V/1997; Schieffer, T.L.; trap: Lindgren Funnel turpentine	[MEM (20) ( : 1)]
186->	United States, Mississippi, Artesia; 28/VII/1929; Hamner, A.L.; Juglandaceae: Carya sp.	[MEM (3) ( : )]
187->	United States, Mississippi, Electric Mills; 13/III/1919; Blackman, M.W.; Juglandaceae: Carya sp.	[USNM (2) (Blackman, M.W. 1920: 449) (Atkinson, T.H. 1989: 327)] [DFEC (1) (Blackman, M.W. 1920: 449) (Atkinson, T.H. 1989: 327)]
188->	United States, Mississippi, Jackson; 26/III/2008; Thomas, B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
189->	United States, Mississippi, M.S.U. Forest T17N, R13E, Sec.11SW; 29/III/1995; Tisdale, R.A.; trap: Lindgren Funnel turpentine + frontalin	[MEM (1) ( : 1)]
190->	United States, Mississippi, McNeill, Miss. State Univ. Exp. Sta.; III/2010; Werle, C.T.; Sampson, B.J.; trap: soda bottle ethanol	[unknown (1) (Werle, C.T., Sampson, B.J, Oliver, J. B. 2012: 3)]
191->	United States, Mississippi, Nicholson; 15/I/1945; Anderson, W.H.; Juglandaceae: Carya sp.	[USNM (1: holotype, female) (Wood, S.L. 1982: 614) (Atkinson, T.H. 1989: 335) (Wood, S.L. 1974: 17)] [USNM (1: paratype(s), female) (Wood, S.L. 1974: 17)]
192->	United States, Mississippi, PEARL; 25/III/2011; Phillips, J.; trap alpha pinene + ethanol	[unknown (12) (EDRR 2011: 0)]
193->	United States, Mississippi, Starkville; 26/VIII/1981; Cross, W.H.; trap: light ultraviolet	[MEM (1) (TH Atkinson-2009, unpublished)]
194->	United States, Mississippi, Starkville; 21/VIII/1981; Cross, W.H.; trap: light ultraviolet	[MEM (1) ( : 1)]
195->	United States, Missouri, Clinton, SE of; 5/V/2004; Nelson, G.H.; Betulaceae: Betula nigra	[FSCA (2) (TH Atkinson-2021, unpublished)]
196->	United States, Missouri, Columbia; 22/V/2014; Doerhoff, R.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
197->	United States, Missouri, Columbia; 9/V/2014; Doerhoff, R.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
198->	United States, Missouri, Columbia; 10/IV/2008; Hook, B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
199->	United States, Missouri, Dent Co.; trap: window trap ethanol	[USNM () (Roling, M.P., Kearby, W.H. 1975: 548) (Atkinson, T.H.; Rabaglia,R.J.; Peck,S.J.; Foltz,J.L. 1991: 157)]
200->	United States, Missouri, Dent Co.; 25/V/1973; Roling, M.P.; trap: window trap ethanol	[SEMC (1) (Atkinson, T.H.; Rabaglia,R.J.; Peck,S.J.; Foltz,J.L. 1991: 157)]
201->	United States, Missouri, Kansas City; 10/IV/2008; Hook, B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
202->	United States, Missouri, Mexico; 9/IV/2008; Hook, B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
203->	United States, Missouri, Platte City; 21/V/2014; Doerhoff, R.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
204->	United States, Missouri, remote location; 22/V/2014; Doerhoff, R.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
205->	United States, Missouri, St. Louis; 12/IV/2008; Hook, B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
206->	United States, Missouri, St. Louis; 17/IV/2008; Walsh, M.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
207->	United States, Missouri, St. Robert; 30/V/2014; Doerhoff, R.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
208->	United States, Missouri, Strafford; 30/V/2014; Doerhoff, R.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
209->	United States, Missouri, Warrenton; 11/IV/2008; Hook, B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
210->	United States, Missouri, Weldon Spring; 19/VI/2014; Doerhoff, R.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
211->	United States, Nebraska, Hwy 81 at Lincoln Creek; 19/V/2007; Turnbow, R.H.	[RHTC (2) (TH Atkinson-2013, unpublished)]
212->	United States, Nebraska, Plattsmouth; 15/IV/2008; Harrell, M.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
213->	United States, New Jersey, Allentown; 2/IV/2008; Zhang, J.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
214->	United States, New Jersey, Cream Ridge; 2/IV/2008; Zhang, J.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
215->	United States, New Jersey, Duke; 20/IV/2009; Golon, D.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
216->	United States, New Jersey, Halka Nursery; 16/IV/2009; Golon, D.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
217->	United States, New Jersey, New Brunswick; 25/III/2008; Zhang, J.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
218->	United States, New Jersey, New Jersey (state)	[unknown () (Wood, S.L. 1982: 426)]
219->	United States, New Jersey, Newton; 16/IV/2010; Oleksak, B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
220->	United States, New Jersey, Pequest	[USNM () (Wood, S.L. 1982: 614) (Atkinson, T.H. 1989: 327)]
221->	United States, New Jersey, Pittstown; 9/IV/2008; Zhang, J.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
222->	United States, New York, Delmar; 7/VI/2012; Cancelliere, J.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
223->	United States, New York, Highland Falls; 20/VI/1915; Schott, P.	[LSAM (1) (TH Atkinson-1988, unpublished)]
224->	United States, New York, Lancaster; 26/VI/2013; Manning, M.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
225->	United States, New York, Lewistown; 13/IV/2011; Cancelliere, J.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
226->	United States, New York, Mount Morris; 16/V/2007; FH field crew; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
227->	United States, New York, New York (state)	[MCZ (1: holotype, female) (Wood, S.L. 1982: 426) (Wood, S.L.; Bright, D.E. 1992: 341)]
228->	United States, New York, Newburgh; 6/VI/2012; Denham, J.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
229->	United States, New York, Pleasantville; 14/IV/2011; Brady, E.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
230->	United States, New York, Rochester; 21/VI/2012; Moskalenko, M.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
231->	United States, New York, Syracuse; 12/V/1916; Stage, H.H.; Juglandaceae: Carya sp.	[USNM (1: holotype) (Blackman, M.W. 1920: 449) (Wood, S.L. 1982: 614) (Blackman, M.W. 1920: 449) (TH Atkinson-2017, unpublished)] [DFEC (1) (Blackman, M.W. 1920: 449) (Blackman, M.W. 1920: 449)] [TAMU (1) (Blackman, M.W. 1920: 449) (Blackman, M.W. 1920: 449)]
232->	United States, New York, Syracuse; 29/XII/1953; Thomas, G.M.; Rosaceae: Malus pumila	[DFEC (50) (TH Atkinson-1988, unpublished)] [TAMU (6) (TH Atkinson-1988, unpublished)]
233->	United States, New York, Syracuse; 6/VI/1972; Jones, W.; Rosaceae: Malus sp.	[UTIC (2) (TH Atkinson-2013, unpublished)] [FSCA (1) (TH Atkinson-2013, unpublished)]
234->	United States, North Carolina, Charlotte; 12/IV/2011; Heath; Lawing; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
235->	United States, North Carolina, Charlotte; 9/V/2013; Lawing; Heath; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
236->	United States, North Carolina, Durham; Juglandaceae: Carya sp.	[Unknown () (Beal, J.A., Massey, C.L. 1945: 764)]
237->	United States, North Carolina, Durham; Juglandaceae: Juglans nigra	[Unknown () (Beal, J.A., Massey, C.L. 1945: 764)]
238->	United States, North Carolina, Greensboro; 14/IV/2011; Heath; Lawing; trap alpha pinene + ethanol	[unknown (5) (EDRR 2011: 0)]
239->	United States, North Carolina, McDowell Co.; 1977; Weber, B.C.; trap: window trap	[unknown (1) (Weber, B.C., MacPherson, J.E. 1991: 46)]
240->	United States, North Carolina, Raleigh; 12/IV/2011; Blaedow, R.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
241->	United States, North Carolina, Raleigh; 27/IV/2013; Oten; Langston; Moan; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
242->	United States, North Carolina, Tryon; Fiske, W.F.; Sapindaceae: Acer sp.	[USNM () (Wood, S.L. 1982: 615) (Atkinson, T.H. 1989: 327)]
243->	United States, North Carolina, Tryon; Fiske, W.F.; Fagaceae: Castanea dentata	[USNM () (Wood, S.L. 1982: 615) (Atkinson, T.H. 1989: 327)]
244->	United States, North Carolina, Tryon; Fiske, W.F.; Juglandaceae: Carya sp.	[USNM () (Wood, S.L. 1982: 615) (Atkinson, T.H. 1989: 327)]
245->	United States, North Carolina, Tryon	[USNM () (Blackman, M.W. 1928: 454) (Blackman, M.W. 1928: 454)]
246->	United States, Ohio, 8-miles north I-71; 17/IV/2007; Nielsen, D.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
247->	United States, Ohio, Ashland; 5/V/2010; Bowers; Zeto; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
248->	United States, Ohio, Ashland; 16/V/2012; Downs; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
249->	United States, Ohio, Columbus; 27/IV/2010; Bowers; Zeto; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
250->	United States, Ohio, Delaware Co.	[Unknown () (Wood, S.L. 1982: 614)]
251->	United States, Ohio, Findlay; 17/V/2012; Downs; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
252->	United States, Ohio, Franklin Co.	[Unknown () (Wood, S.L. 1982: 614)]
253->	United States, Ohio, Lancaster; 29/IV/2010; Bowers; Zeto; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
254->	United States, Ohio, Marietta; 6/V/2010; Bowers; Zeto; trap alpha pinene + ethanol	[unknown (3) (EDRR 2011: 0)]
255->	United States, Ohio, Marietta; 11/V/2012; Downs; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
256->	United States, Ohio, Ohio (state)	[unknown () (Wood, S.L. 1982: 426)]
257->	United States, Ohio, Summit Co; 17/IV/2007	[MSUC (1) (Anonymous, 2010: 0)]
258->	United States, Ohio, Westerville	[Unknown () (Wood, S.L. 1982: 614)]
259->	United States, Oklahoma, Ardmore; 11/V/2011; Hall, B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
260->	United States, Oklahoma, Latimer Co.; VI/1984; Stephan, K.	[TAMU (1) (TH Atkinson-2007, unpublished)]
261->	United States, Oklahoma, Latimer Co.; IX/1985; Stephan, K.	[TAMU (1) (TH Atkinson-2007, unpublished)]
262->	United States, Oklahoma, Latimer Co.; X/1984; Stephan, K.	[TAMU (2) (TH Atkinson-2007, unpublished)]
263->	United States, Oklahoma, Latimer Co.; V/1985; Stephan, K.	[TAMU (1) (TH Atkinson-2007, unpublished)]
264->	United States, Oklahoma, Latimer Co.; VI/1987; Stephan, K.	[TAMU (1) (TH Atkinson-2007, unpublished)]
265->	United States, Oklahoma, Latimer Co.; X/1987; Stephan, K.	[TAMU (1) (TH Atkinson-2007, unpublished)]
266->	United States, Oklahoma, Latimer Co.; III/1990; Stephan, K.	[TAMU (1) (TH Atkinson-2007, unpublished)]
267->	United States, Oklahoma, Latimer Co.; IV/1991; Stephan, K.	[TAMU (3) (TH Atkinson-2007, unpublished)]
268->	United States, Oklahoma, Latimer Co.; V/1991; Stephan, K.	[TAMU (1) (TH Atkinson-2007, unpublished)]
269->	United States, Oklahoma, Latimer Co.; V/1983; Stephan, K.	[CNC (2) (Bright, D.E., Skidmore, R.E. 2002: 514)]
270->	United States, Oklahoma, Latimer Co.; IV/1988; Stephan, K.	[TAMU (1) (TH Atkinson-2019, unpublished)]
271->	United States, Oklahoma, Oklahoma (state)	[unknown (0) (Bright, D.E.; Skidmore, R.S. 1997: 96)]
272->	United States, Oklahoma, Tulsa; 11/V/2011; Hall, B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
273->	United States, Pennsylvania, Allegheny Co.	[CNC (1) (Blackman, M.W. 1920: 449)]
274->	United States, Pennsylvania, Bensalem; 16/IV/2009; Golon, D.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
275->	United States, Pennsylvania, Blain	[USNM () (Wood, S.L. 1982: 614) (Atkinson, T.H. 1989: 327)]
276->	United States, Pennsylvania, Brown; 30/V/2013; Laubscher, B.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
277->	United States, Pennsylvania, Carlisle; 3/IV/2009; Donovall, L; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
278->	United States, Pennsylvania, Carlisle; 3/IV/2009; Donovall, L; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
279->	United States, Pennsylvania, Colley; 25/IV/2011; Stitzer, S.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
280->	United States, Pennsylvania, Franklin; 21/IV/2011; Schmit, D.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
281->	United States, Pennsylvania, Harrisburg	[USNM () (Blackman, M.W. 1928: 454) (Wood, S.L. 1982: 614) (Atkinson, T.H. 1989: 327)]
282->	United States, Pennsylvania, Harrisburg; 2/IV/2009; Spichiger, S.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
283->	United States, Pennsylvania, Liberty; 13/IV/2011; Laubscher, B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
284->	United States, Pennsylvania, Manheim; 20/IV/2011; Schmit, D.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
285->	United States, Pennsylvania, Manheim; 9/IV/2012; Coons, S.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
286->	United States, Pennsylvania, Manheim; 25/IV/2013; Coons, S.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
287->	United States, Pennsylvania, Montoursville; 7/IV/2009; Helbley, K.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
288->	United States, Pennsylvania, Mt. Gretna; 2/IV/2009; Spichiger, S.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
289->	United States, Pennsylvania, Pennsylvania (state)	[unknown () (Wood, S.L. 1982: 426)]
290->	United States, Pennsylvania, Pittsburgh	[Unknown () (Wood, S.L. 1982: 614)]
291->	United States, Pennsylvania, Potter; 11/IV/2011; Wilt, R.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
292->	United States, Pennsylvania, Potter; 10/V/2013; Price, T.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
293->	United States, Pennsylvania, Rush; 28/III/2011; Williams, J.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
294->	United States, Pennsylvania, Shiremanstown; 12/VI/1916	[TAMU (1) (Atkinson, T.H. 1989: 327)] [USNM (3) (Atkinson, T.H. 1989: 327) (Bright, D.E., Skidmore, R.E. 2002: 514)]
295->	United States, Pennsylvania, University Pk.	[Unknown () (Wood, S.L. 1982: 614)]
296->	United States, Pennsylvania, Woodland; 3/IV/2009; Weston, R.; trap alpha pinene + ethanol	[unknown (7) (EDRR 2011: 0)]
297->	United States, Pennsylvania, York; 2/IV/2009; Rebert, S.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
298->	United States, South Carolina, Clemson	[CUAC (1) (Atkinson, T.H. 1989: 327)]
299->	United States, South Carolina, Florence; 12/V/1961; Kirk, V.M.	[USNM (1: holotype, female) (Wood, S.L. 1982: 614)]
300->	United States, South Carolina, Lesslie; 17/III/2010; Reid, L.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
301->	United States, South Carolina, Pageland; 9/III/2010; Reid, L.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
302->	United States, South Carolina, Piedmont; 21/III/2007; Beckham, C.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
303->	United States, South Carolina, Rock Hill; 17/III/2010; Reid, L.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
304->	United States, South Carolina, Rock Hill; 3/IV/2012; Reid, L.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
305->	United States, South Carolina, Savannah River Site; 28/VI/2011; Coyle, D.R.; trap: Lindgren Funnel	[UGCA (1) (TH Atkinson-2011, unpublished)] [USNM (14: paratype(s)) (Wood, S.L. 1971: 8) (TH Atkinson-2022, unpublished)]
306->	United States, Tennessee, Clarksville; 13/V/2009; Jackson, D.; Steele, B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
307->	United States, Tennessee, Kingsport; 27/V/2009; Bingham; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (TH Atkinson-2009, unpublished)]
308->	United States, Tennessee, Kingsport; 13/V/2009; Bingham, S.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
309->	United States, Tennessee, Murfreesboro; 13/V/2009; Strohmeier, C.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
310->	United States, Tennessee, Univ. Tenn. Arboretum; 8/IX/; LaForest, J.M.; Magnoliaceae: Liriodendron tulipifera	[ECUT (1) (TH Atkinson-2000, unpublished)]
311->	United States, Texas, Bentsen-Rio Grande St. Pk..; 15/IV/1977; Turnbow, R.H.; Cannabaceae: Celtis lindheimeri	[TAMU (1) (Atkinson, T.H.; Rabaglia,R.J.; Peck,S.J.; Foltz,J.L. 1991: 158)]
312->	United States, Texas, Big Slough Wldlf. Area; 9/V/1986; Anderson, R.	[CMNC (1) (TH Atkinson-1991, unpublished)]
313->	United States, Texas, Clarkesville; 25/III/2010; Prater, E.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
314->	United States, Texas, Conroe; 15/III/2007; Kramer, M.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
315->	United States, Texas, Hemphill, 8 mi E; 27/IV/2008; Riley, E.G.; Raber, B.; Heffern, D.J.; trap: Lindgren Funnel	[TAMU (1) (TH Atkinson-2013, unpublished) (DE Bright-2011, unpublished)]
316->	United States, Texas, Hemphill, 8.8 mi NE, Mill Creek Cove; 31/III/2008; Riley, E.G.; et al; trap: flight intercept	[EGRC (1) (TH Atkinson-2008, unpublished)]
317->	United States, Texas, Hemphill, 9 mi E; 4/IV/1989; Anderson, R.S.; Morris, R.; trap: malaise	[TAMU (1) (TH Atkinson-2009, unpublished)]
318->	United States, Texas, Hempstead; 1/XII/2006; Smith, J.E.; Bogran, C.; trap: Lindgren Funnel	[TAMU (4) (TH Atkinson-2012, unpublished)]
319->	United States, Texas, Hempstead; 1/XII/2006; Smith, J.E.; Bogran, C.; trap: bottle trap	[TAMU (3) (TH Atkinson-2012, unpublished)]
320->	United States, Texas, Hempstead; 2/XI/2006; Smith, J.E.; Bogran, C.; trap: bottle trap	[TAMU (3) (TH Atkinson-2012, unpublished)]
321->	United States, Texas, Hempstead; 2/XI/2006; Smith, J.E.; Bogran, C.; trap: Lindgren Funnel	[TAMU (2) (TH Atkinson-2012, unpublished)]
322->	United States, Texas, Hempstead; 22/II/2007; Smith, J.E.; Bogran, C.; trap: Lindgren Funnel	[TAMU (1) (TH Atkinson-2012, unpublished)]
323->	United States, Texas, Houston; 20/III/2011; Marquez, S.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
324->	United States, Texas, Humble; 11/IV/2014; McArthur, T.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
325->	United States, Texas, Jasper; 24/IV/2013; Horne, L.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
326->	United States, Texas, LaRue; 23/X/2013; Atkinson, T.H.; trap: alpha-scents 4 vane ethanol	[UTIC (1) (TH Atkinson-2014, unpublished)]
327->	United States, Texas, Lufkin; 19/III/2007; Clarke; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
328->	United States, Texas, Lufkin; 21/IV/2014; Clarke; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
329->	United States, Texas, Magnolia; 30/IV/2014; McArthur, T.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
330->	United States, Texas, Mt. Pleasant; 21/IV/2013; Semrau, A.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
331->	United States, Texas, Paige; 19/III/2007; Brooks, B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
332->	United States, Texas, Sam Houston NF, jct hwy 149 & 1791; 11/IV/1998; Riley, E.G.; trap: Lindgren Funnel	[TAMU (1) (TH Atkinson-2012, unpublished)]
333->	United States, Texas, TAMU Farm; 13/VIII/1982; Turnbow, R.H.; Juglandaceae: Carya illinoensis	[TAMU (1) (Atkinson, T.H.; Rabaglia,R.J.; Peck,S.J.; Foltz,J.L. 1991: 158)] [CNC (1) (Bright, D.E., Skidmore, R.E. 2002: 514)]
334->	United States, Texas, Tivoli, 7 mi SW Hwy 35; 4/IV/1998; Riley, E.G.	[TAMU (1) (TH Atkinson-2012, unpublished)]
335->	United States, Texas, Waterwood; 21/V/2014; Russell, S.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
336->	United States, Texas, Willis; 25/I/2007; Smith, J.E.; Bogran, C.; trap: Lindgren Funnel	[TAMU (1) (TH Atkinson-2012, unpublished)]
337->	United States, Texas, Willis; 22/II/2007; Smith, J.E.; Bogran, C.; trap: Lindgren Funnel	[TAMU (1) (TH Atkinson-2019, unpublished)]
338->	United States, Texas, Woodville; 11/VI/2007	[MSUC (1) (Anonymous, 2010: 0)]
339->	United States, Texas, Woodville; 19/XI/2007	[MSUC (1) (Anonymous, 2010: 0)]
340->	United States, Vermont, Castleton; 16/VI/1989; Weems, H.V.	[FSCA (1) (TH Atkinson-2007, unpublished)]
341->	United States, Virginia, Newport News; 21/V/2013; Johnson; Ansicar; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
342->	United States, Virginia, Richmond; 28/III/2008; Hubbard, H.G.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
343->	United States, Virginia, Sandston; 17/V/2013; Hubbard, H.G.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
344->	United States, Virginia, Shenandoah; 21/V/2013; McGehee, J.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
345->	United States, Virginia, Virginia (state)	[unknown () (Wood, S.L. 1982: 614)]
346->	United States, West Virginia, Arthur; 24/III/2009; Long, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
347->	United States, West Virginia, Keyser; 23/III/2009; Hern, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
348->	United States, West Virginia, Triadelphis; 30/III/2009; Geiger, W.; Pownall, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
349->	United States, Wisconsin, Sheboygan; 15/IV/2009; Murphy, R.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
UNK
350->	, no state / prov. data; Rosaceae: Crataegus sp.	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 341)]
351->	, no state / prov. data; Rosaceae: Sorbus sp.	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 341)]