Bark and Ambrosia Beetles of

SYNONYMY

• Gnathotrichus denticulatus Blackman 1931. J. Wash. Acad. Sci. 21: 270

DISTRIBUTION.

Numbers in parentheses after each geographic unit are the number of distinct collection events in the database for that unit. For exotic species generally only countries are listed for localities outside the New World. For further information on published sources of distribution, check the REFERENCES section.

• Central America (32)
  Belize (1): Orange Walk (1); Costa Rica (2): Alajuela (1), Puntarenas (1); El Salvador (1): La Libertad (1); Guatemala (10): Alta Verapaz (1), Escuintla (2), Huehuetenango (1), Peten (1), Quetzaltenango (1), San Marcos (1), Suchitepequez (2), Zacapa (1); Honduras (5): Atlántida (3), Comayagua (1), Francisco Morazán (1); Nicaragua (1): Jinotega (1); Panamá (12): Canal Zone (1), Chiriquí (1), Coclé (1), Panamá (8), Veraguas (1);
• Caribbean (20)
  Cuba (3): Santiago de Cuba (2), unknown (1); Jamaica (5): no state / prov. data (4), unknown (1); Lesser Antilles (6): Dominica (2), Guadeloupe (1), Martinique (1), St. Lucia (1), St. Vincent (1); Rep. Dominicana (6): no state / prov. data (2), Dajabon (1), Duarte (1), Pedernales (1), Sanchez Ramirez (1);
• Eurasia (1)
  Italy (1): no state / prov. data (1);
• North America (160)
  (2): no state / prov. data (2); México (41): Chiapas (1), Chihuahua (1), Distrito Federal (3), Durango (2), Guanajuato (1), Guerrero (2), México (3), Michoacán (1), Morelos (5), Nayarit (2), Nuevo León (1), Oaxaca (1), Puebla (5), San Luís Potosí (2), Tamaulipas (1), unknown (1), Veracruz (9); United States (117): no state / prov. data (2), Arizona (32), California (1), Colorado (27), Florida (26), Georgia (2), Louisiana (1), Montana (2), Nevada (1), New Mexico (6), North Carolina (1), South Carolina (1), South Dakota (4), Texas (6), Utah (4), Washington (1);
• South America (21)
  (1): no state / prov. data (1); Argentina (3): Tucumán (2), unknown (1); Bolivia (1): unknown (1); Brazil (1): Amazonas (1); Chile (1): Arica y Parinacota (I) (1); Colombia (3): Narino (1), Tolima (1), Valle del Cauca (1); Ecuador (1): Galápagos (1); Fr. Guiana (1): Cayenne (1); Paraguay (1): no state / prov. data (1); Perú (3): Huánuco (2), unknown (1); Trinidad and Tobago (3): Trinidad (3); Venezuela (2): D.C. (2);

HOSTS

Numbers in parentheses after each host family, genus, or collection method are the number of distinct collection events in the database for that host or method.

• No host information (151);
• beating (3): (3): no species data (3);
• Fagaceae (1): Quercus (1): chapmani (1);
• Lauraceae (23): Nectandra (3): coriacea (3); Persea (20): no species data (2), americana (12), borbonia (4), sp. (2);
• litter (3): (2): no species data (1), wet pluviselva (1); berlese (1): upper montane cloud fores (1);
• Myrtaceae (1): Myrica (1): cerifera (1);
• Pinaceae (16): Abies (1): concolor (1); Picea (1): engelmanni (1); Pinus (14): montezumae (1), ponderosa (9), sp. (3), strobiformis (1);
• Poaceae (2): Zea (2): maiz (1), mays (1);
• trap (34) (6): alpha pinene + ethanol (6); 'tampa colgante' (1): no details (1); flight intercept (2): no details (2); funnel (1): ethanol (1); light (10): no details (3), mercury vapor + UV (1), ultraviolet (6); Lindgren Funnel (10): no details (1), alpha pinene + ethanol (3), ethanol (5), exotic ips lure (1); malaise (3): no details (3); yellow pan trap (1): no details (1);

REFERENCES

The following are important recent monographs, catalogs, and supplements to catalogs that refer to this species. The specific page on which the reference is made is shown in pink at the end of the reference. In the case of Wood (1982) and Wood & Bright (1992) clicking on the reference page will link to a digital version of the work in question.

• Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6:1-1356. [1163]. (data capture complete)
• Wood, S.L., Bright,D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index. Great Basin Nat. Mem. 13:1-1553 (vol. A, B). [1035]

The following are references from which host and distribution data have been input into the database. If one of the above monographs or catalogs also appears in this list, it means that most relevant collection event data have been included.

1. . . .
2. EDRR 2011. USFS Early Detection Rapid Response Database. http://www.fs.fed.us/invasivespecies/earlydetection.shtml/.
3. EDRR 2014. USFS Early Detection Rapid Response Database. http://www.fs.fed.us/invasivespecies/earlydetection.shtml/.
4. Gast,S.J.; Furniss,M.M.; Johnson,J.B.; Ivie,M.A. 1989. List of Montana Scolytidae (Coleoptera) and notes on new records. Great Basin Nat. 49(3): 381-386.
5. University of Arizona Insect Collection 2016. SCAN: Symbiota Collections of Arthropods Network. http://symbiota4.acis.ufl.edu/scan/portal/collections/misc/collprofiles.php?collid=11.
6. Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6: 1-1356.
7. Wood, S.L.; Bright, D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index (Volumes A,B). Great Basin Nat. Mem. 13: 1-1553.

Maps automatically open at the center of the plotted points and the scale is set to encompass all map points. Maps can be resized (scale bar at upper left) and the center moved (place cursor over map and drag) to see other parts of the distribution of the species. Clicking on a map marker will pull up collection event data and a literature citation if present. If any errors are found, please refer to the "series code" which is a unique identifier for a database record in any communications). Coordinates have not been entered for all collection records. Localities outside the New World are not plotted, even though they are listed in the distribution summary and in the table of records.

No.	Collection data	Publication
Central America
1->	Belize, Orange Walk, Rio Bravo Cons. Area, Well Trail; 15/IV/1995; Kovarik, P.W.; trap: yellow pan trap	[CAS (2) (TH Atkinson-2012, unpublished)]
2->	Costa Rica, Alajuela, Monteverde; 3/VI/1984; Riley, E.G.; et al	[LSAM (2) (TH Atkinson-1989, unpublished)]
3->	Costa Rica, Puntarenas, Est. Biol. Las Alturas, 2 km NE Alturas; 20/VI/1998; Anderson, R.; litter: berlese upper montane cloud fores	[FSCA (1) (TH Atkinson-2007, unpublished)]
4->	El Salvador, La Libertad, San Andrés	[Unknown () (Wood, S.L. 1982: 246)]
5->	Guatemala, Alta Verapaz, Senahú	[BMNH (1) (Blandford, W.F.H. 1905: 135)]
6->	Guatemala, Escuintla, El Zapote	[BMNH (1) (Blandford, W.F.H. 1905: 135)]
7->	Guatemala, Escuintla, Pantaleon	[BMNH (1) (Blandford, W.F.H. 1905: 135)]
8->	Guatemala, Huehuetenango, Res. Nat. Protegida Yal Unin Yul Witz; 13/VII/2018; Zack, R.S.; trap: light	[WSU (1) (TH Atkinson-2021, unpublished)]
9->	Guatemala, Peten, Río Machaquila; 19/X/2017; Zack, R.S.; trap: light	[WSU (2) (TH Atkinson-2021, unpublished)]
10->	Guatemala, Quetzaltenango, Cerro Zunil	[BMNH (1) (Blandford, W.F.H. 1905: 135)]
11->	Guatemala, San Marcos, El Tumbador	[BMNH (1) (Blandford, W.F.H. 1905: 135)]
12->	Guatemala, Suchitepequez, Finca Los Tarrales; 16/VII/2018; Zack, R.S.; trap: light	[WSU (1) (TH Atkinson-2021, unpublished)]
13->	Guatemala, Suchitepequez, San Isidro	[BMNH (1) (Blandford, W.F.H. 1905: 135)]
14->	Guatemala, Zacapa, Mirandilla	[BMNH (1) (Blandford, W.F.H. 1905: 135)]
15->	Honduras, Atlántida, Lancetilla; 30/V/1996; Andrews, F.G.; Hardy, A.; trap: light ultraviolet	[CSCA (1) (TH Atkinson-2021, unpublished)]
16->	Honduras, Atlántida, Parque Nal. Pico Bonito, El Portillo; 1/VII/2000; Reyes, R.	[EAPZ (1) (TH Atkinson-2013, unpublished)]
17->	Honduras, Atlántida, Parque Nal. Pico Bonito, Est. CURLA; 18/VII/2001; Turnbow, R.H.	[RHTC (1) (TH Atkinson-2008, unpublished)]
18->	Honduras, Comayagua, La Libertad, 4 km N; 18/VII/1977; O'Brien, C.W.; O'Brien, L.B.; Marshall, G.B.	[CWOB () (Atkinson, T.H.; Equihua M., A. 1988: )]
19->	Honduras, Francisco Morazán, El Zamorano; 23/II/2001; Uclés, D.; Suazo, C.	[EAPZ (1) (TH Atkinson-2013, unpublished)]
20->	Nicaragua, Jinotega, Parque Nacional Cerro Saslaya; 15/V/2011; LLAMA Project; beating	[SEMC (1) (TH Atkinson-2020, unpublished)]
21->	Panamá, Canal Zone, Canal Zone	[Unknown () (Wood, S.L. 1982: 246)]
22->	Panamá, Chiriquí, Volcán	[NMW (1: holotype) (Wood, S.L. 1982: 246)]
23->	Panamá, Coclé, La Campana	[Unknown () (Wood, S.L. 1982: 246)]
24->	Panamá, Panamá, Cerro Azul; 13/II/1999; Turnbow, R.H.	[RHTC (1) (TH Atkinson-2008, unpublished)]
25->	Panamá, Panamá, Cerro Azul; 3/VI/1993; Andrews, F.G.; Gilbert, A.J.	[CSCA (1) (TH Atkinson-2021, unpublished)]
26->	Panamá, Panamá, El Llano-Carti Rd.; 9/I/1994; Wappes, J.E.	[TAMU (1) (TH Atkinson-2006, unpublished)]
27->	Panamá, Panamá, El Llano-Carti Rd.; 13/II/1999; Turnbow, R.H.	[RHTC (1) (TH Atkinson-2008, unpublished)]
28->	Panamá, Panamá, El Llano-Carti Rd., km 6-8; 6/VI/1994; Andrews, F.G.; Gilbert, A.J.	[CSCA (1) (TH Atkinson-2021, unpublished)]
29->	Panamá, Panamá, El Llano-Carti Rd., km 9; 17/II/1992; Stockwell, H.P.	[TAMU (1) (TH Atkinson-2006, unpublished)]
30->	Panamá, Panamá, El Llano-Carti Rd., km 9; 16/V/1984; Stockwell, H.P.	[TAMU (1) (TH Atkinson-2006, unpublished)]
31->	Panamá, Panamá, El Llano-Carti Rd., km 9; 21/V/1993; Riley, E.G.	[EGRC (1) (TH Atkinson-2012, unpublished)]
32->	Panamá, Veraguas, Cerro Tute, 8 km W Santa Fe; 24/VII/1999; Wooley, J.B.; trap: malaise	[TAMU (1) (TH Atkinson-2012, unpublished)]
Caribbean
33->	Cuba, Santiago de Cuba, PN Gran Piedra; 27/I/2012; Anderson, R.S.; litter wet pluviselva	[CMNC (1) (Bright, D.E. 2019: 29)]
34->	Cuba, Santiago de Cuba, PN Gran Piedra; 27/I/2012; Anderson, R.S.; litter
35->	Cuba, unknown, Cuba (country)	[ZUMH (1: syntype(s)) (Wood, S.L. 1982: 246)]
36->	Jamaica, no state / prov. data, Hardwar Gap	[CNC (2) (Bright, D.E. 2019: 29)]
37->	Jamaica, no state / prov. data, Kellits; 4/VIII/1974; Peck, S.B.; Peck, J.	[SBP (1) (Bright, D.E. 2019: 29)]
38->	Jamaica, no state / prov. data, Mandeville	[SBP (2) (Bright, D.E. 2019: 29)]
39->	Jamaica, no state / prov. data, Portland Gap; 1/VIII/1974; Peck, S.B.	[SBP (1) (Bright, D.E. 2019: 990)]
40->	Jamaica, unknown, Jamaica (country)	[unknown (0) (Bright, D.E.; Skidmore, R.S. 1997: 44)]
41->	Lesser Antilles, Dominica, Roseau	[USNM (2) (Bright, D.E. 2019: 29)]
42->	Lesser Antilles, Dominica, Springfield Estate; 31/V/1995; Eden; Stieber; Wooley, J.B.	[TAMU (1) (TH Atkinson-2006, unpublished)]
43->	Lesser Antilles, Guadeloupe, Trois Rivieres	[NMW (1: holotype) (Wood, S.L. 1982: 246) (Eggers, H. 1940: 136) (Wood, S.L.; Bright, D.E. 1992: 213)]
44->	Lesser Antilles, Martinique, Grand Riviére; 21/VII/2001; Roguet; Marquet	[CNCI (1) (Bright, D.E. 2019: 990)]
45->	Lesser Antilles, St. Lucia, Barre de L'isle; 2009	[WIBF (1) (Bright, D.E. 2019: 29)]
46->	Lesser Antilles, St. Vincent, Hermitage Forest; 15/VIII/2006; Peck, S.B.; Peck, J.; trap: flight intercept	[SBP (1) (Bright, D.E. 2019: 29)]
47->	Rep. Dominicana, no state / prov. data, La Vega, .7 km S Jarabacoa; 24/V/1992; Thomas, M.C.	[FSCA (1) (TH Atkinson-2007, unpublished)]
48->	Rep. Dominicana, no state / prov. data, Mt Diego de Ocampo; 1/VII/1938; Darlington	[MCZ (1) (TH Atkinson-1988, unpublished)]
49->	Rep. Dominicana, Dajabon, Mariano Cestero; 14/VIII/1980; Norrbom, A.	[CMNH (1) (Bright, D.E. 2019: 29)]
50->	Rep. Dominicana, Duarte, Reserva LomaQuita Espuela, Canelo; 6/IV/2004; Young, C.; Davidson, R.; Rawlins, J.; trap: light ultraviolet	[CMNH (5) (Bright, D.E. 2019: 29)]
51->	Rep. Dominicana, Pedernales, Cabo Rojo, 24 km N; 11/VII/1993; Sikes, D.; Rosenfeld, R.; beating	[WIBF (1) (Bright, D.E. 2019: 29)]
52->	Rep. Dominicana, Sanchez Ramirez, Sanchez; 1/VII/1938; Darlington	[MCZ (1) (TH Atkinson-1988, unpublished)]
Eurasia
53->	Italy, no state / prov. data, Genoa	[IEGG (1: holotype) (Wood, S.L. 1982: 246)]
North America
54->	, no state / prov. data; Pinaceae: Pinus strobiformis	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 1035)]
55->	, no state / prov. data; Pinaceae: Abies concolor	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 1035)]
56->	México, Chiapas, Teopisca, 17 mi SE; 3/VI/1969; Howden, H.F.	[CNC (1) (Bright, D.E.; Skidmore, R.S. 1997: 44) (Bright, D.E., Skidmore, R.E. 2002: 517)]
57->	México, Chihuahua, Ojinaga	[unknown () (Wood, S.L. 1982: 1163)]
58->	México, Distrito Federal, D.F.	[Unknown () (Schedl, K.E. 1940: )]
59->	México, Distrito Federal, Distrito Federal; Lauraceae: Persea americana	[Unknown () (Wood, S.L. 1982: 246)]
60->	México, Distrito Federal, El Guarda	[unknown () (Wood, S.L. 1982: 1163)]
61->	México, Durango, Durango	[unknown () (Wood, S.L. 1982: 1163)]
62->	México, Durango, El Salto	[unknown () (Wood, S.L. 1982: 1163)]
63->	México, Guanajuato, Celaya; 7/X/2014; Marín J., A.; Lauraceae: Persea americana	[CEAM (34) (TH Atkinson-2020, unpublished)]
64->	México, Guerrero, Acapulco	[unknown () (Wood, S.L. 1982: 1163)]
65->	México, Guerrero, Chilpancingo; 12/XII/1958; Rivera, J.	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 230)]
66->	México, México, Carr. Toluca-Zitácuaro, km 54; 30/X/1980; Atkinson, T.H.; Equihua M., A.; Pinaceae: Pinus montezumae	[CEAM (2) (Atkinson, T.H.; Equihua M., A. 1985: 361) (TH Atkinson-2014, unpublished)] [TAMU (2) (Atkinson, T.H.; Equihua M., A. 1985: 361)]
67->	México, México, Chapingo; Lauraceae: Persea sp.	[Unknown () (Wood, S.L. 1982: 246)]
68->	México, México, Malinalco; 20/XII/1956; Moreno, H.	[CEAM (2) (Atkinson, T.H.; Equihua M., A. 1985: 230) (TH Atkinson-2020, unpublished)]
69->	México, Michoacán, Carápan	[unknown () (Wood, S.L. 1982: 1163)]
70->	México, Morelos, Carr. Tetela-Hueyapan, km 2; 19/V/1984; Saucedo C., E.; Lauraceae: Persea americana	[CEAM (1) (Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986: 20) (TH Atkinson-2020, unpublished)]
71->	México, Morelos, Jumiltepec; 28/I/1982; Burgos S., A.; Lauraceae: Persea americana	[UAEM () (Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986: 20)]
72->	México, Morelos, Oacalco; Lauraceae: Persea americana	[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 48)]
73->	México, Morelos, Tepoztlan; 23/IV/1982; Saucedo C., E.; Burgos S., A.; Lauraceae: Persea americana	[CEAM (1) (Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986: 20) (TH Atkinson-2020, unpublished)]
74->	México, Morelos, Tlalnepantla; 4/VII/1989; Equihua M., A.; Lauraceae: Persea americana	[CEAM (22) (Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986: 20) (TH Atkinson-2020, unpublished)]
75->	México, Nayarit, 8 km S Santiago Ferry	[Unknown () (Wood, S.L. 1982: 246)]
76->	México, Nayarit, Xalisco; II/2019; Lauraceae: Persea americana	[UAEM (1) (Avila, N.D.; Ríos V., C.; Cambero C., J.; Burgos Solorio, A.; Dominguez M., J.L.; Estrada V., M.O. 2021: 320)]
77->	México, Nuevo León, Linares, 16 mi W Hwy 85; 21/VII/1982; O'Brien, C.W.; O'Brien, L.B.; Wibmer, G.	[FSCA (1) (TH Atkinson-2007, unpublished)]
78->	México, Oaxaca, Comaltepec; 14/VIII/2014; Atkinson, T.H.; Burgos S., A.; Lauraceae: Persea americana	[UTIC (1) (TH Atkinson-2014, unpublished)]
79->	México, Puebla, Atlixco; 24/X/1980; Bautista M., N.; Lauraceae: Persea americana	[CEAM (13) (Atkinson, T.H.; Equihua M., A. 1985: 230) (TH Atkinson-2020, unpublished)]
80->	México, Puebla, Texmelucán	[unknown () (Wood, S.L. 1982: 1163)]
81->	México, Puebla, Teziutlan, 24 km E	[Unknown () (Wood, S.L. 1982: 246)]
82->	México, Puebla, Tiaguismanalco; 19/I/1983; Atkinson, T.H.; Lauraceae: Persea americana	[CEAM (6) (Atkinson, T.H.; Equihua M., A. 1985: 230) (TH Atkinson-2020, unpublished)]
83->	México, Puebla, Zacapoaxtla, 27 mi N.; 10/VI/1983; O'Brien, C.W.; O'Brien, L.B.; Marshall, G.B.	[FSCA (4) (TH Atkinson-2007, unpublished)]
84->	México, San Luís Potosí, San Luís Potosí	[unknown () (Wood, S.L. 1982: 1163)]
85->	México, San Luís Potosí, San Luís Potosí (estado)	[TAMU (1) (TH Atkinson-, unpublished)]
86->	México, Tamaulipas, Bocatoma, 7 km SSE G. Farías; 25/III/1978; Riley, E.G.	[EGRC (1) (TH Atkinson-2011, unpublished)]
87->	México, unknown, Mexico (country); I/2002	[TAMU (3) (TH Atkinson-2008, unpublished)]
88->	México, Veracruz, Est. Biol. Los Tuxtlas; 11/XI/1989; Colin, J.L.; Rojas, H.; trap: 'tampa colgante'	[CNIN (1) (TH Atkinson-2018, unpublished)]
89->	México, Veracruz, Los Tuxtlas Biol. Sta.; 14/XI/1989; Colin, J.L.; Rojas, H.	[FSCA (1) (TH Atkinson-2007, unpublished)]
90->	México, Veracruz, Los Tuxtlas Biol. Sta.; 16/X/1989; Colin, J.L.	[FSCA (1) (TH Atkinson-2007, unpublished)]
91->	México, Veracruz, Los Tuxtlas Biol. Sta.; 13/I/1986; Andrade, D.	[FSCA (1) (TH Atkinson-2007, unpublished)]
92->	México, Veracruz, Los Tuxtlas Biol. Sta.; 21/X/1985; Andrade, D.	[FSCA (1) (TH Atkinson-2007, unpublished)]
93->	México, Veracruz, Los Tuxtlas Biol. Sta.; 1/IX/1988; Mariño, E.; Mejorada, E.	[FSCA (1) (TH Atkinson-2007, unpublished)]
94->	México, Veracruz, Los Tuxtlas Biol. Sta.; 22/V/1983; O'Brien, C.W.; O'Brien, L.B.; Marshall, G.B.	[CAS (1) (TH Atkinson-2007, unpublished)]
95->	México, Veracruz, Misantla, 1 mi. S	[TAMU (3) (TH Atkinson-, unpublished)]
96->	México, Veracruz, Naranjos, 7.3 mi E	[TAMU (4) (TH Atkinson-, unpublished)]
97->	United States, no state / prov. data; Pinaceae: Picea engelmanni	[unknown (0) (Wood, S.L. 1982: 1163)]
98->	United States, no state / prov. data, 'Carolina'	[ZMUC (1: syntype(s)) (Wood, S.L. 1982: 246) (Wood, S.L. 2007: 99)]
99->	United States, Arizona, Chiricahua Mtns.	[unknown () (Wood, S.L. 1982: 1163)]
100->	United States, Arizona, Flagstaff	[unknown () (Wood, S.L. 1982: 1163)]
101->	United States, Arizona, Flagstaff; 2/IX/1985; Beal, R.S.; Pinaceae: Pinus ponderosa	[CPMAB (1) (TH Atkinson-2021, unpublished)]
102->	United States, Arizona, Flagstaff (near); 10/V/1977; Sanderson, M.W.	[UAIC (15) (University of Arizona Insect Collection 2016: 0)]
103->	United States, Arizona, FS 300, 0.2 mi W Hwy 260; 15/X/2011; Warner, W.B.; Pinaceae: Pinus ponderosa	[WBWC (3) (TH Atkinson-2013, unpublished)] [UTIC (2) (TH Atkinson-2013, unpublished)]
104->	United States, Arizona, Ft. Valley For. Turp. Area; 5/1959; Nutting, W.L.; Pinaceae: Pinus ponderosa	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
105->	United States, Arizona, Ft. Valley For. Turp. Area; 5/IX/1959; Nutting, W.L.; Pinaceae: Pinus ponderosa	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
106->	United States, Arizona, Jerome	[unknown () (Wood, S.L. 1982: 1163)]
107->	United States, Arizona, Kaibab N.F.	[unknown () (Wood, S.L. 1982: 1163)]
108->	United States, Arizona, Mt. Graham; 18/V/1969; Stephan, K.	[FSCA (1) (DE Bright-2010, unpublished)]
109->	United States, Arizona, Onion Saddle, 1 mi S, Chiricahua Mtns; 5/V/1988; Andrews, F.G.; Eichlin, T.; Pinaceae: Pinus sp.	[CSCA (1) (TH Atkinson-2014, unpublished)]
110->	United States, Arizona, Paradise	[unknown () (Wood, S.L. 1982: 1163)]
111->	United States, Arizona, Pinal Mtns.; 22/VIII/1935; Parker, F.H.	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
112->	United States, Arizona, Prescott; 22/I/1959; Ostmark, H.E.; Pinaceae: Pinus ponderosa	[FSCA (20) (WH Anderson-1969, unpublished) (TH Atkinson-1987, unpublished)]
113->	United States, Arizona, Prescott N.F.	[unknown () (Wood, S.L. 1982: 1163)]
114->	United States, Arizona, Rincon Mtns.	[unknown () (Wood, S.L. 1982: 1163)]
115->	United States, Arizona, Rustler Park, 0.8 mi NW; 26/VII/1989; Andrews, F.G.; Eichlin, T.	[CSCA (2) (TH Atkinson-2014, unpublished)]
116->	United States, Arizona, Rustler Park, Chiricahua Mtns.; 17/VI/1973; Chandler, D.S.	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
117->	United States, Arizona, Rustlers Park; 14/VIII/1970; McCleve, S.; Pinaceae: Pinus sp.	[UAIC (4) (University of Arizona Insect Collection 2016: 0)]
118->	United States, Arizona, Rustlers Park; 31/VIII/1974; McCleve, S.; Pinaceae: Pinus sp.	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
119->	United States, Arizona, San Francisco Peak	[unknown () (Wood, S.L. 1982: 1163)]
120->	United States, Arizona, Sitgreaves N.F.	[unknown () (Wood, S.L. 1982: 1163)]
121->	United States, Arizona, Sta. Catalina Mtns.	[unknown () (Wood, S.L. 1982: 1163)]
122->	United States, Arizona, Sta. Catalina Mtns.; 19/V/1968; Stephan, K.	[FSCA (1) (DE Bright-2010, unpublished)]
123->	United States, Arizona, Sta. Catalina Mtns.; 7/IX/1968; Stephan, K.	[FSCA (1) (DE Bright-2010, unpublished)]
124->	United States, Arizona, Sta. Catalina Mtns.; 12/IV/1969; Stephan, K.	[FSCA (1) (DE Bright-2010, unpublished)]
125->	United States, Arizona, Strayhorse Campground; 30/V/1970; Stephan, K.	[FSCA (1) (DE Bright-2010, unpublished)]
126->	United States, Arizona, Wheatfields Lake; 6/VI/1976; Stephan, K.; trap: light ultraviolet	[FSCA (1) (TH Atkinson-1987, unpublished)]
127->	United States, Arizona, White Mtns.; 9/VII/1936; Parker, F.H.	[UAIC (4) (University of Arizona Insect Collection 2016: 0)]
128->	United States, Arizona, White River	[unknown () (Wood, S.L. 1982: 1163)]
129->	United States, Arizona, Williams	[unknown () (Wood, S.L. 1982: 1163)]
130->	United States, Arizona, Wupatki Natl. Mon.; 14/VIII/1978; Peters, J.; trap: light ultraviolet	[CPMAB (1) (TH Atkinson-2021, unpublished)]
131->	United States, California, Chiquito Basin	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
132->	United States, Colorado, Allenspark; 14/VI/2011; Daniel, Z.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
133->	United States, Colorado, Allenspark; 17/V/2012; Daniel, Z.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
134->	United States, Colorado, Blackhawk; 3/V/2012; Daniel, Z.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
135->	United States, Colorado, Boulder; 13/VI/2011; Daniel, Z.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
136->	United States, Colorado, Caribou; 17/V/2012; Daniel, Z.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
137->	United States, Colorado, Carter Lake; 4/V/2012; Rowley, T.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
138->	United States, Colorado, Colorado N. F.	[unknown () (Wood, S.L. 1982: 1163)]
139->	United States, Colorado, Colorado Springs	[unknown () (Wood, S.L. 1982: 1163)]
140->	United States, Colorado, Estes Park	[unknown () (Wood, S.L. 1982: 1163)]
141->	United States, Colorado, Estes Park; 13/VI/2011; Rowley, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
142->	United States, Colorado, Estes Park; 4/V/2012; Rowley, T.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
143->	United States, Colorado, Evergreen	[unknown () (Wood, S.L. 1982: 1163)]
144->	United States, Colorado, Fort Collins; 4/V/2012; Rowley, T.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
145->	United States, Colorado, Fort Collins; 21/V/2012; Rowley, T.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
146->	United States, Colorado, Ft. Collins	[unknown () (Wood, S.L. 1982: 1163)]
147->	United States, Colorado, Gilpin; 16/VI/2011; Daniel, Z.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
148->	United States, Colorado, Glen Haven	[unknown () (Wood, S.L. 1982: 1163)]
149->	United States, Colorado, Gunnison	[unknown () (Wood, S.L. 1982: 1163)]
150->	United States, Colorado, Livermore; 14/VI/2011; Rowley, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
151->	United States, Colorado, Longmont; 2/V/2012; Daniel, Z.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
152->	United States, Colorado, Nederland; 14/VI/2011; Daniel, Z.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
153->	United States, Colorado, Norwood	[unknown () (Wood, S.L. 1982: 1163)]
154->	United States, Colorado, Ouray	[unknown () (Wood, S.L. 1982: 1163)]
155->	United States, Colorado, Stove Praire; 21/V/2012; Rowley, T.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
156->	United States, Colorado, Uncompahgre N.F.	[unknown () (Wood, S.L. 1982: 1163)]
157->	United States, Colorado, Waldo Canyon; 9/VIII/1916; Edmonston, W.D.; Pinaceae: Pinus ponderosa	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
158->	United States, Colorado, Waldo Canyon	[unknown () (Wood, S.L. 1982: 1163)]
159->	United States, Florida, Amelia Island	[unknown () (Wood, S.L. 1982: 246)]
160->	United States, Florida, Archbold Biol. Sta.; 1978; Deyrup, M.A.; et al	[ABS (21) (MA Deyrup-, unpublished) (TH Atkinson-1987, unpublished)]
161->	United States, Florida, Archbold Biol. Sta.; 7/II/1986; Deyrup, M.A.; Myrtaceae: Myrica cerifera	[ABS (1) (MA Deyrup-, unpublished) (TH Atkinson-1987, unpublished)]
162->	United States, Florida, Archbold Biol. Sta.; 1/IV/1985; Deyrup, M.A.; Lauraceae: Persea borbonia	[ABS (1) (MA Deyrup-, unpublished) (TH Atkinson-1987, unpublished)]
163->	United States, Florida, Archbold Biol. Sta.; 31/I/1986; Deyrup, M.A.; Fagaceae: Quercus chapmani	[ABS (1) (MA Deyrup-, unpublished) (TH Atkinson-1987, unpublished)]
164->	United States, Florida, Archbold Biol. Sta.; 16/IX/1990; Turnbow, R.H.	[RHTC (1) (TH Atkinson-1993, unpublished)]
165->	United States, Florida, Brevard Co.	[unknown () (Wood, S.L. 1982: 246)]
166->	United States, Florida, Chekika St. Rec. Area; 12/X/1990; Anderson, R.; beating	[CMNC (1) (TH Atkinson-1991, unpublished)]
167->	United States, Florida, Chekika St. Rec. Area, 50 km SW Miami; 15/XI/1985; Peck, S.; Peck, J.	[CMNC (2) (TH Atkinson-1989, unpublished)]
168->	United States, Florida, Coronado Beach; 1939; Frost, C.A.	[MCZ (2) (TH Atkinson-1988, unpublished)]
169->	United States, Florida, Deering Estate Pk.; 1989; Peck, S.; Peck, J.; trap: flight intercept	[CMNC (1) (TH Atkinson-1989, unpublished)]
170->	United States, Florida, Deering Estate Pk.; 26/VIII/; Peck, S.; Peck, J.; trap: malaise	[CMNC (4) (TH Atkinson-1991, unpublished)]
171->	United States, Florida, Edgewater; 28/II/1939; Frost, C.A.	[MCZ (3) (TH Atkinson-1988, unpublished)]
172->	United States, Florida, Ft. Lauderdale; 14/IX/1978; Schimmel, R.; Lauraceae: Persea	[FSCA (4) (TH Atkinson-1988, unpublished)]
173->	United States, Florida, Ft. Lauderdale; 9/IX/1953; Link, O.D.; Lauraceae: Nectandra coriacea	[FSCA (100) (TH Atkinson-1988, unpublished)] [CEAM (4) (TH Atkinson-2020, unpublished)]
174->	United States, Florida, Ft. Lauderdale	[FSCA (1) (MC Thomas-2012, downloaded from Bugguide)]
175->	United States, Florida, Gainesville	[unknown () (Wood, S.L. 1982: 246)]
176->	United States, Florida, Little River; 2/IX/1918; Lauraceae: Nectandra coriacea	[unknown () (Wood, S.L. 1982: 246)]
177->	United States, Florida, Marion Co.	[MCZ (1) (TH Atkinson-1988, unpublished)]
178->	United States, Florida, Miami; Lauraceae: Persea borbonia	[unknown () (Wood, S.L. 1982: 246)]
179->	United States, Florida, Miami; 15/VIII/1975; Dowling, C.F.; Lauraceae: Nectandra coriacea	[FSCA (5) (TH Atkinson-1988, unpublished)]
180->	United States, Florida, Monroe Station; 28/VIII/1965; Suter, W.S.	[ABS (1) (MA Deyrup-, unpublished) (TH Atkinson-1987, unpublished)]
181->	United States, Florida, Satsuma Heights; Lauraceae: Persea	[unknown () (Wood, S.L. 1982: 246)]
182->	United States, Florida, St. Lucie	[USNM (1: holotype) (Wood, S.L. 1982: 246) (TH Atkinson-2018, unpublished) (Wood, S.L. 2007: 99)]
183->	United States, Florida, Vero Beach (South Beach); 23/IX/1976; Frank, J.H.; trap: light ultraviolet	[CEAM (6) (TH Atkinson-2020, unpublished)]
184->	United States, Florida, West Palm Beach; 9/IV/1925	[MCZ (1) (TH Atkinson-1988, unpublished)]
185->	United States, Georgia, Savannah; Lauraceae: Persea borbonia	[Unknown () (Wood, S.L. 1982: 246)]
186->	United States, Georgia, St. Simons Isl; 13/VII/1931; Quirsfeld	[MCZ (1) (TH Atkinson-1988, unpublished)]
187->	United States, Louisiana, Abita Creek Pres.; 3/XI/1999; Carlton, C.E.; Prowell, D.; trap: malaise	[LSAM (1) (TH Atkinson-2013, unpublished)]
188->	United States, Montana, Cow Creek, Custer N.F.; 1/VI/1988; Furniss, M.M.; Gast, S.J.; Pinaceae: Pinus ponderosa	[WFBM (19) (Gast,S.J.; Furniss,M.M.; Johnson,J.B.; Ivie,M.A. 1989: 384)]
189->	United States, Montana, Lantis Springs Campground; 2/VI/1988; Furniss, M.M.; Gast, S.J.; Pinaceae: Pinus ponderosa	[WFBM (1) (Gast,S.J.; Furniss,M.M.; Johnson,J.B.; Ivie,M.A. 1989: 384)]
190->	United States, Nevada, Kyle Canyon; 19/VI/1969; Stephan, K.	[FSCA (2) (DE Bright-2010, unpublished)]
191->	United States, New Mexico, Capitan	[unknown () (Wood, S.L. 1982: 1163)]
192->	United States, New Mexico, Cloudcroft	[USNM (1: holotype, female) (Wood, S.L. 1982: 1163) (TH Atkinson-2017, unpublished)]
193->	United States, New Mexico, Magdalena	[unknown () (Wood, S.L. 1982: 1163)]
194->	United States, New Mexico, Meek	[unknown () (Wood, S.L. 1982: 1163)]
195->	United States, New Mexico, Oak Grove Campground, Lincoln NF; 29/V/1998; Riley, E.G.; trap: light ultraviolet	[EGRC (3) (TH Atkinson-2008, unpublished)]
196->	United States, New Mexico, Vermejo	[unknown () (Wood, S.L. 1982: 1163)]
197->	United States, North Carolina, North Carolina (state)	[Unknown () (Wood, S.L. 1982: 246)]
198->	United States, South Carolina, Folly Beach; Lauraceae: Persea sp.	[Unknown () (Wood, S.L. 1982: 246)]
199->	United States, South Dakota, Black Hills	[unknown () (Wood, S.L. 1982: 1163)]
200->	United States, South Dakota, Cheyenne Crossing	[unknown () (Wood, S.L. 1982: 1163)]
201->	United States, South Dakota, Elmore	[unknown () (Wood, S.L. 1982: 1163)]
202->	United States, South Dakota, Hill City	[unknown () (Wood, S.L. 1982: 1163)]
203->	United States, Texas, Angelina NF, 7 mi NE Rockland; 19/IX/1966; Clarke; Menard; Riley, E.G.; trap: Lindgren Funnel	[TAMU (1) (EG Riley-2008, unpublished) (TH Atkinson-2008, unpublished)]
204->	United States, Texas, Corpus Christi; 12/VII/1985; Mercer, J.	[TAMU (1) (TH Atkinson-1987, unpublished)]
205->	United States, Texas, Davis Mtns.	[unknown () (Wood, S.L. 1982: 1163)]
206->	United States, Texas, Goose Isl. St. Pk.; 30/XI/1975; Clark, W.E.	[TAMU (36) (TH Atkinson-1987, unpublished)]
207->	United States, Texas, Goose Isl. St. Pk.; 8/I/1992; Jones, S.; Jones, G.; Lauraceae: Persea borbonia	[TAMU (5) (TH Atkinson-2003, unpublished)]
208->	United States, Texas, Weslaco; Lauraceae: Persea americana	[Unknown () (Wood, S.L. 1982: 246)]
209->	United States, Utah, Dixie N.F.	[unknown () (Wood, S.L. 1982: 1163)]
210->	United States, Utah, Elk Park, Ashley N.F.	[unknown () (Wood, S.L. 1982: 1163)]
211->	United States, Utah, Long Hollow, Dixie N.F.	[unknown () (Wood, S.L. 1982: 1163)]
212->	United States, Utah, Panguitch	[unknown () (Wood, S.L. 1982: 1163)]
213->	United States, Washington, Kooskooskie; 13/V/1949; Chastain, G.; Pinaceae: Pinus ponderosa	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
South America
214->	, no state / prov. data, 'South America'	[USNM (1: lectotype) (Wood, S.L. 1982: 246) (TH Atkinson-2018, unpublished) (Wood, S.L.; Bright, D.E. 1992: 213) (Anderson, W.H.; Anderson, D.M. 1971: 36) (Wood, S.L. 2007: 99)]
215->	Argentina, Tucumán, Sauce Huacho; 4/IV/2017; Cordoba S., P.; Mendoza A., E.	[UTIC (1) (TH Atkinson-2017, unpublished)] [IFML (6) (Córdoba, S.P.; Atkinson, T.H. 2018: 376)]
216->	Argentina, Tucumán, Sauce Huacho; XI/2016; Cordoba, S.P.; trap: funnel ethanol	[IFML (1) (Córdoba, S.P.; Mendoza, E.A.; Atkinson, T.H.; Manzo, V.V. 2021: 122)]
217->	Argentina, unknown, Argentina (country)	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 213)]
218->	Bolivia, unknown, Bolivia (country)	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 213)]
219->	Brazil, Amazonas, Rio Solimões; 1/X/1982; Adis, J.	[unknown (1) (Wood, S.L. 2007: 99)]
220->	Chile, Arica y Parinacota (I), Tarapacá; XI/1996; Jimenez, J.	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 213) (Wood, S.L. 2007: 99)]
221->	Colombia, Narino, Pasto; 3/VII/1957; Uniparro, A.; Poaceae: Zea mays	[unknown (1) (Wood, S.L. 2007: 99)]
222->	Colombia, Tolima, Fresno; XII/1939; Carmona B., C.	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 213) (Wood, S.L. 2007: 99)]
223->	Colombia, Valle del Cauca, Palmira; 23/VII/1958; Bravo, C.	[unknown (1) (Wood, S.L. 2007: 99)]
224->	Ecuador, Galápagos, Isla Sta. Cruz	[unknown (0) (Bright, D.E.; Skidmore, R.S. 1997: 44)]
225->	Fr. Guiana, Cayenne, Cayenne, 67 km S on N2 towards Regina; 6/VI/1986; Riley, E.G.; Rider, D.A.; trap: light mercury vapor + UV	[LSAM (1) (TH Atkinson-2013, unpublished)]
226->	Paraguay, no state / prov. data, Paraguay (country); Freiburg	[ZMBN (1) (TH Atkinson-2021, unpublished)]
227->	Perú, Huánuco, Tingo Maria; 15/XII/1954; Schlinger, E.I.; Ross, E.S.	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 213) (Wood, S.L. 2007: 99) (Smith, S.M.; Petrov, A.V.; Cognato, A.I. 2017: 78)]
228->	Perú, Huánuco, Tingo Maria; 17/XII/1946; Kuschel	[NMW (1) (Schedl, K.E. 1960: 6)]
229->	Perú, unknown, Perú (country); 30/III/1966; Poaceae: Zea maiz	[CSCA (10) (TH Atkinson-2016, unpublished)]
230->	Trinidad and Tobago, Trinidad, Ariho Valley; 18/IV/1935; Weber, N.A.	[MCZ (13) (TH Atkinson-1988, unpublished)]
231->	Trinidad and Tobago, Trinidad, Talparo; 5/VII/1989; Dozier, H.L.	[FSCA (1) (TH Atkinson-2007, unpublished)]
232->	Trinidad and Tobago, Trinidad, Trinidad (island)	[unknown (0) (Bright, D.E.; Skidmore, R.S. 1997: 44)]
233->	Venezuela, D.C., Caracas	[NMW (1: holotype) (Wood, S.L. 1982: 246)]
234->	Venezuela, D.C., Caracas; 23/XI/1955	[unknown (1) (Wood, S.L. 2007: 99)]