Bark and Ambrosia Beetles of

SYNONYMY

• Ambrosiophilus osumiensis (Murayama 1934) Annot. Zool. Jap. 14(3): 292
• Xyleborus osumiensis Murayama 1934. Annot. Zool. Jap. 14(3): 292
• Euwallacea metanepotulus (Eggers 1939) Arb. Morphol. Taxon. Entomol. 6: 119
• Ambrosiophilus metanepotulus (Eggers 1939) Arb. Morphol. Taxon. Entomol. 6: 119
• Xyleborus metanepotulus Eggers 1939. Arb. Morphol. Taxon. Entomol. 6: 119
• Ambrosiodmus nodulosus (Eggers 1941) Entomol. Blätter 37: 223
• Ambrosiophilus nodulosus (Eggers 1941) Entomol. Blätter 37: 223
• Premnobius nodulosus (Eggers 1941) Entomol. Blätter 37: 223
• Xyleborus nodulosus Eggers 1941. Entomol. Blätter 37: 223
• Xyleborus pernodulus Schedl 1957. Ann. Mus. Roy. Afr. Cent. Sci. Zool. Ser. 8 56: 85
• Ambrosiophilus hunanensis (Browne 1983) Entomol. Monthly Mag. 119: 33
• Xyleborus hunanensis Browne 1983. Entomol. Monthly Mag. 119: 33
• Ambrosiophilus peregrinus Smith & Cognato 2015. Coleop. Bull. 69(2): 216

DISTRIBUTION.

Numbers in parentheses after each geographic unit are the number of distinct collection events in the database for that unit. For exotic species generally only countries are listed for localities outside the New World. For further information on published sources of distribution, check the REFERENCES section.

• Asia (20)
  China (15): Anhui (1), Chongqing (4), Fukien (1), Guangxi (3), Guizhou (1), Hunan (1), Jiangxi (1), Sichuan (1), Taiwan (1), Yunnan (1); Vietnam (5): Cao Bang (1), Hoa Binh (1), Ninh Binh (1), Thua Thien-Hue (1), Yen Bai (1);
• Eurasia (2)
  China (1): Taiwan (1); Japan (1): no state / prov. data (1);
• North America (5)
  United States (5): Georgia (5);

HOSTS

Numbers in parentheses after each host family, genus, or collection method are the number of distinct collection events in the database for that host or method.

• No host information (14);
• Anacardiaceae (1): Pistacia (1): chinensis (1);
• Aquifoliaceae (1): Ilex (1): oldhami (1);
• Fagaceae (2): Quercus (2): griffithii (1), sp. (1);
• Lauraceae (2): (1): no species data (1); Cinnamomum (1): cassia (1);
• Moraceae (1): Broussonetia (1): papyrifera (1);
• Oleaceae (1): Ligustrum (1): lucidum (1);
• trap (5) Lindgren Funnel (3): no details (2), Ips trilure (1); walnut twig beetle (2): no details (2);

REFERENCES

The following are important recent monographs, catalogs, and supplements to catalogs that refer to this species. The specific page on which the reference is made is shown in pink at the end of the reference. In the case of Wood (1982) and Wood & Bright (1992) clicking on the reference page will link to a digital version of the work in question.

• Wood, S.L., Bright,D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index. Great Basin Nat. Mem. 13:1-1553 (vol. A, B). [758]

The following are references from which host and distribution data have been input into the database. If one of the above monographs or catalogs also appears in this list, it means that most relevant collection event data have been included.

1. Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2017. The ambrosia beetle Ambrosiophilus peregrinus Smith and Cognato, introduced to the USA, is Ambrosiophilus nodulosus (Eggers), new combination (Coleoptera: Curculionidae: Scolytinae). Coleopterists Bull. 71(3): 552-553.
2. Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2018. New synonymy, new combinations and other taxonomic changes in Japanese xyleborine ambrosia beetles (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4251(3): 391-403.
3. Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2020. A monograph of the Xyleborini (Coleoptera, Curculionidae, Scolytinae) of the Indochinese Peninsula (except Malaysia) and southern China. ZooKeys 983: 1-442.
4. Smith, S.M.; Cognato, A.I. 2015. Ambrosiophilus peregrinus Smith and Cognato, New Species (Coleoptera: Curculionidae: Scolytinae), an Exotic Ambrosia Beetle Discovered in Georgia, USA . Coleopterists Bull. 69(2): 213-220.
5. Smith, S.M.; Rabaglia, R.J.; Beaver, R.A.; Thu, P.Q.; Cognato, A.I. 2018. Attraction of Ambrosia Beetles (Coleoptera: Curculionidae: Scolytinae: Xyleborini) to Semiochemicals in Vietnam, with New Records and a New Species. Coleopterists Bull. 72(4): 838-844.
6. Wood, S.L.; Bright, D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index (Volumes A,B). Great Basin Nat. Mem. 13: 1-1553.

Maps automatically open at the center of the plotted points and the scale is set to encompass all map points. Maps can be resized (scale bar at upper left) and the center moved (place cursor over map and drag) to see other parts of the distribution of the species. Clicking on a map marker will pull up collection event data and a literature citation if present. If any errors are found, please refer to the "series code" which is a unique identifier for a database record in any communications). Coordinates have not been entered for all collection records. Localities outside the New World are not plotted, even though they are listed in the distribution summary and in the table of records.

No.	Collection data	Publication
Asia
1->	China, Anhui, Chuxian; Anacardiaceae: Pistacia chinensis	[USNM (4) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2020: 65)]
2->	China, Chongqing, Mt. Simian; 5/VI/2015; Ly-Jia, T.-S.	[RAB (2) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2017: 552)]
3->	China, Chongqing, Nan Shan; 20/VIII/2015; Wang, J.-G.; Ly-Jia, T.-S.	[RAB (4) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2017: 552)]
4->	China, Chongqing, Pengshui; 11/VIII/2016; Ly-Jia, T.-S.	[RAB (2) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2017: 552)]
5->	China, Chongqing, Youyang; 14/V/2016; Ly-Jia, T.-S.	[RAB (1) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2017: 552)]
6->	China, Fukien, Kuatun; 4/IV/1938; Klapperich, L.J.	[ZFMK (1: holotype, female) (Wood, S.L.; Bright, D.E. 1992: 677) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2020: 65) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2017: 552)]
7->	China, Guangxi, Malu; Lauraceae: Cinnamomum cassia	[UFFE (1) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2020: 65)]
8->	China, Guangxi, Shangsi; Moraceae: Broussonetia papyrifera	[UFFE (1) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2020: 65)]
9->	China, Guangxi, Shiwandashan; Fagaceae: Quercus griffithii	[UFFE (1) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2020: 65)]
10->	China, Guizhou, Guizhou (province)	[unknown (1) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2020: 65)]
11->	China, Hunan, Changsha; 15/VI/1977; Huang, F.S.	[IZCAS (1: holotype, female) (Wood, S.L.; Bright, D.E. 1992: 743) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2020: 65)]
12->	China, Jiangxi, Ganzhou; Oleaceae: Ligustrum lucidum	[RAB (1) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2020: 65)]
13->	China, Sichuan, Mt. Emei; 19/VIII/2016; Ly-Jia, T.-S.	[RAB (1) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2017: 552)]
14->	China, Taiwan, Taiwan (province)	[unknown (1) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2020: 65)]
15->	China, Yunnan, Xishuangbanna, Sanchahe Nat. Res.; 29/V/2008; Cognato, A.I.; Fagaceae: Quercus sp.	[MSUC (1) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2020: 65)]
16->	Vietnam, Cao Bang, coordinates only	[MSUC (1) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2020: 65)]
17->	Vietnam, Hoa Binh, Hoa Binh (province)	[USNM (1) (Smith, S.M.; Rabaglia, R.J.; Beaver, R.A.; Thu, P.Q.; Cognato, A.I. 2018: 843)]
18->	Vietnam, Ninh Binh, coordinates only; Lauraceae	[MSUC (2) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2020: 65)]
19->	Vietnam, Thua Thien-Hue, Bach Ma N.P.; 15/II/2017; Cognato, A.I.; Hoang, T.A.	[MSUC (1 , female) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2020: 65)]
20->	Vietnam, Yen Bai, Yen Bai (province)	[RJRC (1) (Smith, S.M.; Rabaglia, R.J.; Beaver, R.A.; Thu, P.Q.; Cognato, A.I. 2018: 843)]
Eurasia
21->	China, Taiwan, Hoorin Kwarenko; 25/XII/1933; Chujo, M.	[NMNST (1: holotype, female) (Wood, S.L.; Bright, D.E. 1992: 692) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2018: 393) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2020: 65)]
22->	Japan, no state / prov. data, Nagao Forest, Osumi Peninsula; Aquifoliaceae: Ilex oldhami	[USNM (1: holotype, female) (Wood, S.L.; Bright, D.E. 1992: 758) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2018: 393)]
North America
23->	United States, Georgia, Ball Ground, 11.3 km SE; 9/IX/2013; VanAtta, C.; trap: Lindgren Funnel Ips trilure	[UGCA (1) (Smith, S.M.; Cognato, A.I. 2015: 216)]
24->	United States, Georgia, Chatsworth, 17.3 km SSE; 10/VII/2013; VanAtta, C.; trap: Lindgren Funnel	[USNM (1: holotype, female) (Smith, S.M.; Cognato, A.I. 2015: 216) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2020: 65)] [CMNH (1: paratype(s), female) (Smith, S.M.; Cognato, A.I. 2015: 216)] [MSUC (5: paratype(s), female) (Smith, S.M.; Cognato, A.I. 2015: 216) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2020: 65)]
25->	United States, Georgia, Chatsworth, 17.3 km SSE; 13/VI/2013; VanAtta, C.; trap: Lindgren Funnel	[USNM (2: paratype(s), female) (Smith, S.M.; Cognato, A.I. 2015: 216)] [CMNH (2: paratype(s), female) (Smith, S.M.; Cognato, A.I. 2015: 216)] [MSUC (4: paratype(s), female) (Smith, S.M.; Cognato, A.I. 2015: 216)]
26->	United States, Georgia, Chatsworth, near; 30/IX/2014; Sullivan, J.; trap: walnut twig beetle	[UGCA (1: paratype(s), female) (Smith, S.M.; Cognato, A.I. 2015: 216)]
27->	United States, Georgia, Cohutta, near; 16/X/2014; Sullivan, J.; trap: walnut twig beetle	[UGCA (1: paratype(s), female) (Smith, S.M.; Cognato, A.I. 2015: 216)]